''Apatoraptor'' ("Apatè robber") is a genus of

caenagnathid Caenagnathidae is a family of bird-like maniraptoran theropod dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and close relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathids ...

dinosaur Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...

which contains a single species, ''A. pennatus''. The only known specimen was discovered in the

Campanian The Campanian is the fifth of six ages of the Late Cretaceous Epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campani ...

-age

Horseshoe Canyon Formation The Horseshoe Canyon Formation is a stratigraphic unit of the Western Canada Sedimentary Basin in southwestern Alberta. It takes its name from Horseshoe Canyon, an area of badlands near Drumheller. The Horseshoe Canyon Formation is part of the ...

Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest Ter ...

Discovery

In 1993, at

Drumheller Drumheller is a town on the Red Deer River in the badlands of east-central Alberta, Canada. It is northeast of Calgary and south of Stettler. The Drumheller portion of the Red Deer River valley, often referred to as Dinosaur Valley, has a ...

, three kilometres west of the ''

Royal Tyrrell Museum of Palaeontology The Royal Tyrrell Museum of Palaeontology (RTMP, and often referred to as the Royal Tyrrell Museum) is a palaeontology museum and research facility in Drumheller, Alberta, Canada. The museum was named in honour of Joseph Burr Tyrrell, and is situ ...

'', a skeleton was found of a theropod. As it was initially identified as some unimportant ornithomimid specimen, preparation only started in 2002 and was almost immediately discontinued when no skull was thought to be present, the fossil again being covered in plaster. In 2008, research was resumed, and only then was it discovered that the fossil represented a species new to science. In 2016, the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

''Apatoraptor pennatus'' was named and described by Gregory F. Funston and Philip John Currie. The generic name is derived from the Greek goddess of deceit, Apatè, and

Latin Latin (, or , ) is a classical language belonging to the Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through the power ...

''raptor'', "robber", in reference to the specimen hiding its true identity for many years. The specific name means "feathered" in Latin, referring to the find of quill knobs on the

ulna The ulna (''pl''. ulnae or ulnas) is a long bone found in the forearm that stretches from the elbow to the smallest finger, and when in anatomical position, is found on the medial side of the forearm. That is, the ulna is on the same side of t ...

, showing the animal had wings. The

holotype A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of seve ...

TMP 1993.051.0001 was found in a layer of the Horsethief Member of the Horseshoe Canyon Formation dating from the Campanian. It consists of a partial skeleton with skull. The fossil was largely found articulated. It contains a palatine bone, the lower jaws, a

hyoid The hyoid bone (lingual bone or tongue-bone) () is a horseshoe-shaped bone situated in the anterior midline of the neck between the chin and the thyroid cartilage. At rest, it lies between the base of the mandible and the third cervical vertebra. ...

, a complete series of neck and back vertebrae, ribs, the right shoulder girdle, the right arm, a sternum, belly ribs, a right ilium, a right thighbone, a right shinbone and a right calfbone. It was further investigated by means of a

CAT-scan A computed tomography scan (CT scan; formerly called computed axial tomography scan or CAT scan) is a medical imaging technique used to obtain detailed internal images of the body. The personnel that perform CT scans are called radiographers ...

Classification

''Apatoraptor'' was, within the

Oviraptorosauria Oviraptorosaurs ("egg thief lizards") are a group of feathered maniraptoran dinosaurs from the Cretaceous Period of what are now Asia and North America. They are distinct for their characteristically short, beaked, parrot-like skulls, with or w ...

, placed in the

Caenagnathidae Caenagnathidae is a family of bird-like maniraptoran theropod dinosaurs from the Cretaceous of North America and Asia. They are a member of the Oviraptorosauria, and close relatives of the Oviraptoridae. Like other oviraptorosaurs, caenagnathi ...

, in a derived position as a possible

sister species In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

of ''

Elmisaurus ''Elmisaurus'' (meaning "foot sole lizard") is an extinct genus of caenagnathid dinosaur from the Late Cretaceous Nemegt Formation of Mongolia. It was a theropod belonging to the Oviraptorosauria. Discovery In 1970, a paleontological Polish- ...

''.

References

{{Taxonbar, from=Q23807302 Caenagnathids Campanian life Late Cretaceous dinosaurs of North America Fossils of Canada Paleontology in Alberta Fossil taxa described in 2016 Taxa named by Philip J. Currie

Discovery

Classification

See also

References