''Ancylus'' is a

genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...

of very small, freshwater, air-breathing

limpets Limpets are a group of aquatic snails that exhibit a conical shell shape (patelliform) and a strong, muscular foot. Limpets are members of the class Gastropoda, but are polyphyletic, meaning the various groups called "limpets" descended inde ...

. They are aquatic pulmonate gastropod

mollusk Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is es ...

s in the tribe

Ancylini Ancylini is a tribe of small, freshwater, air-breathing limpets, aquatic pulmonate gastropod mollusks in the family Planorbidae, the ram's horn snails and their allies. This tribe used to be treated as a family; the current taxonomic placement ...

within the family

Planorbidae Planorbidae, common name the ramshorn snails or ram's horn snails, is a family of air-breathing freshwater snails, aquatic pulmonate gastropod molluscs. Unlike most molluscs, the blood of ram's horn snails contains iron-based hemoglobin inste ...

, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. (2014). Ancylus O. F. Müller, 1773. Accessed through: World Register of Marine Species at http://www.marinespecies.org/aphia.php?p=taxdetails&id=559196 on 2014-11-15 The genus is known throughout the

Cenozoic The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configu ...

Species

Species in the genus ''Ancylus'' include: * ''

Ancylus aduncus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' A. A. Gould, 1847"Species in genus ''Ancylus'' (n=6)"
AnimalBase, accessed 7 July 2012. * † ''

Ancylus alutae ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. (20 ...

'' Jekelius, 1932 * ''

Ancylus ashangiensis ''Ancylus ashangiensis'' is a species of very small, air-breathing, freshwater snail or limpet. It is an aquatic pulmonate gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates ...

'' Brown, 1965 * ''

Ancylus benoitianus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Bourguignat, 1862 * † ''

Ancylus bourgeoisi ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Deshayes, 1863 * † ''

Ancylus boyeri ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Noulet, 1867 * † ''

Ancylus braunii ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Dunker, 1853 * ''

Ancylus capuloides ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic animal, aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rose ...

'' Jan in Porro, 1838 * † ''

Ancylus cestasensis ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Peyrot, 1932 * † ''

Ancylus depressus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Deshayes, 1824 * † ''

Ancylus dumasi ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Fontannes, 1884 * † ''

Ancylus elegans ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' J. de C. Sowerby, 1826 * ''

Ancylus expansilabris ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Clessin, 1881 * ''

Ancylus fluviatilis The river limpet (''Ancylus fluviatilis'') is a species of very small, freshwater, air-breathing limpet, an aquatic pulmonate gastropod mollusk in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies. ...

'' Müller, 1774 - the

type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specimen( ...

* † ''

Ancylus hungaricus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Brusina, 1902 * ''

Ancylus lapicidus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Hubendick, 1960 * † ''

Ancylus lyelli ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Maillard, 1892 * † ''

Ancylus moravicus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Rzehak, 1893 * † ''

Ancylus ninghaiensis ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Youluo, 1978 * † ''

Ancylus parmophorus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' de Stefani, 1880 * ''

Ancylus pileolus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. (20 ...

'' * ''

Ancylus recurvus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. (20 ...

'' Martens, 1873 * ''

Ancylus regularis ''Ancylus regularis'' is a species of very small, freshwater, air-breathing limpet, an aquatic pulmonate gastropod mollusk in the tribe Ancylini Ancylini is a tribe of small, freshwater, air-breathing limpets, aquatic pulmonate gastropod ...

'' Brown, 1973Van Damme D. (2010)
''Ancylus regularis''
In: IUCN 2012. IUCN Red List of Threatened Species. Version 2012.1. . Downloaded on 7 July 2012. * ''

Ancylus rupicola ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Boubée, 1832 * † ''

Ancylus saucatsensis ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Peyrot, 1932 (?) * ''

Ancylus scalariformis ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. (20 ...

'' Stankovic & Radoman, 1953 * † ''

Ancylus serbicus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Brusina, 1893 * ''

Ancylus striatus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Quoy &

Gaimard Joseph Paul Gaimard (31 January 1793 – 10 December 1858) was a French naval surgeon A naval surgeon, or less commonly ship's doctor, is the person responsible for the health of the ship's company aboard a warship. The term appears often ...

, 1834 * ''

Ancylus subcircularis ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Clessin, 1880 * † ''

Ancylus subcostatus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Benoist, 1873 * ''

Ancylus tapirulus ''Ancylus'' is a genus of very small, freshwater, air-breathing limpets. They are aquatic pulmonate gastropod mollusks in the tribe Ancylini within the family Planorbidae, the ram's horn snails and their allies.Bouchet, P.; Rosenberg, G. ( ...

'' Polinski, 1929 ;Synonyms: * † ''Ancylus croaticus'' Brusina, 1902: synonym of † ''

Acroloxus croaticus ''Acroloxus'' is a genus of very small, air-breathing, freshwater snails, or more precisely limpets, aquatic pulmonate gastropods in the family Acroloxidae. Species Species in the genus ''Acroloxus'' include: * ''Acroloxus coloradensis'' J. H ...

'' (Brusina, 1902) * † ''Ancylus decussatus'' Reuss in Reuss & Meyer, 1849: synonym of †''

Acroloxus decussatus ''Acroloxus'' is a genus of very small, air-breathing, freshwater snails, or more precisely limpets, aquatic pulmonate gastropods in the family Acroloxidae. Species Species in the genus ''Acroloxus'' include: * ''Acroloxus coloradensis'' J. H ...

'' (Reuss in Reuss & Meyer, 1849) * † ''Ancylus deperditolacustris'' Gottschick, 1911: synonym of † ''

Acroloxus deperditolacustris ''Acroloxus'' is a genus of very small, air-breathing, freshwater snails, or more precisely limpets, Aquatic animal, aquatic pulmonate gastropods in the family Acroloxidae. Species Species in the genus ''Acroloxus'' include: * ''Acroloxus colora ...

'' (Gottschick, 1911) * † ''Ancylus deperditus'' (Desmarest, 1814): synonym of † ''

Ferrissia deperdita ''Ferrissia'' is a genus of small, air-breathing freshwater limpets, aquatic gastropod mollusks in the family Planorbidae, the ram's horn snails and their allies. Species Species within the genus ''Ferrissia'' include: * '' Ferrissia baconi ...

'' (Desmarest, 1814) * ''Ancylus drouetianus'' Bourguignat, 1862: synonym of ''

Williamia gussoni ''Williamia'' is a genus of small sea snails or false limpets, marine pulmonate gastropod molluscs in the family Siphonariidae, the false (air-breathing) limpets.Bouchet, P.; Gofas, S. (2011). Williamia Monterosato, 1884. Accessed through: World ...

'' (Costa O. G., 1829) * ''Ancylus fuscus'' C. B. Adams, 1840: synonym of ''

Laevapex fuscus ''Laevapex fuscus'', the dusky ancylid, is a species of small, freshwater, air-breathing limpet, an aquatic pulmonate gastropod mollusc or micromollusc in the family Planorbidae, the ram's horn snails and their allies. Geographic distrib ...

'' (C. B. Adams, 1840) * ''Ancylus gibbosus'' Bourguignat, 1862: synonym of ''

'' (Costa O. G., 1829) * ''Ancylus gussonii'' O. G. Costa, 1829: synonym of ''

Williamia gussonii ''Williamia'' is a genus of small sea snails or false limpets, marine pulmonate gastropod molluscs in the family Siphonariidae, the false (air-breathing) limpets.Bouchet, P.; Gofas, S. (2011). Williamia Monterosato, 1884. Accessed through: Wo ...

'' (O. G. Costa, 1829) * † ''Ancylus illyricus'' Neumayr, 1880: synonym of †''

Ferrissia illyrica ''Ferrissia'' is a genus of small, air-breathing freshwater limpets, aquatic gastropod mollusks in the family Planorbidae, the ram's horn snails and their allies. Species Species within the genus ''Ferrissia'' include: * ''Ferrissia baconi'' ( ...

'' (Neumayr, 1880) * † ''Ancylus involutus'' Pavlović, 1903: synonym of † ''

Acroloxus involutus ''Acroloxus'' is a genus of very small, air-breathing, freshwater snails, or more precisely limpets, aquatic pulmonate gastropods in the family Acroloxidae. Species Species in the genus ''Acroloxus'' include: * ''Acroloxus coloradensis'' J. H ...

'' (Pavlović, 1903) * † ''Ancylus michaudi'' Locard, 1878: synonym of † ''

Acroloxus michaudi ''Acroloxus'' is a genus of very small, air-breathing, freshwater snails, or more precisely limpets, aquatic pulmonate gastropods in the family Acroloxidae. Species Species in the genus ''Acroloxus'' include: * ''Acroloxus coloradensis'' J. H ...

'' (Locard, 1878) * † ''Ancylus neumayri'' Fontannes, 1881: synonym of †''

Ferrissia neumayri ''Ferrissia'' is a genus of small, air-breathing freshwater limpets, aquatic gastropod mollusks in the family Planorbidae, the ram's horn snails and their allies. Species Species within the genus ''Ferrissia'' include: * '' Ferrissia baconi ...

'' (Fontannes, 1881) * ''Ancylus patelloides'' I. Lea, 1856: synonym of ''

Lanx patelloides A ''lanx'' (Latin for dish) was a large ancient Roman serving platter.Lysons, Daniel & Lysons, Samuel"Antiquities: British and Roman" Magna Britannia vol 5, 1817, pp. CCIII-CCXVIII. Accessed November 26, 2007. Particularly ornamented ones were used ...

'' (I. Lea, 1856) * † ''Ancylus pontileviensis'' Morgan, 1920: synonym of † ''

Ferrissia pontileviensis ''Ferrissia'' is a genus of small, air-breathing freshwater limpets, aquatic gastropod mollusks in the family Planorbidae, the ram's horn snails and their allies. Species Species within the genus ''Ferrissia'' include: * ''Ferrissia baconi'' ( ...

'' (Morgan, 1920) * † ''Ancylus wittmanni'' Schlickum, 1964: synonym of † ''

Ferrissia wittmanni ''Ferrissia'' is a genus of small, air-breathing freshwater limpets, aquatic gastropod mollusks in the family Planorbidae, the ram's horn snails and their allies. Species Species within the genus ''Ferrissia'' include: * '' Ferrissia baconi ...

'' (Schlickum, 1964)

Cladogram

A cladogram based on sequences of mitochondrial

16S ribosomal RNA 16 S ribosomal RNA (or 16 S rRNA) is the RNA component of the 30S subunit of a prokaryotic ribosome ( SSU rRNA). It binds to the Shine-Dalgarno sequence and provides most of the SSU structure. The genes coding for it are referred to as 16S r ...

and cytochrome-c oxidase I (COI) genes showing phylogenic relations of ''Ancylus'' by Albrecht et al. (2006):Albrecht C., Trajanovski S., Kuhn K., Streit B. & Wilke T. (2006). "Rapid evolution of an ancient lake species flock: Freshwater limpets (Gastropoda: Ancylidae) in the Balkan Lake Ohrid". ''

Organisms Diversity & Evolution In biology, an organism () is any living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy into groups such as multicellular animals, plants, and fungi ...

'' 6(4): 294-307. .

Species

Cladogram

References

Further reading