''Anaxyrus'', containing the North American toads, is a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

true toad A true toad is any member of the family Bufonidae, in the order Anura (frogs and toads). This is the only family of anurans in which all members are known as toads, although some may be called frogs (such as harlequin frogs). The bufonids now ...

s in the

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

Bufonidae {{Cat main, Toad This category Category, plural categories, may refer to: General uses *Classification, the general act of allocating things to classes/categories Philosophy * Category of being * ''Categories'' (Aristotle) * Category (Kant) * C ...

. The genus is

endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...

North North is one of the four compass points or cardinal directions. It is the opposite of south and is perpendicular to east and west. ''North'' is a noun, adjective, or adverb indicating Direction (geometry), direction or geography. Etymology T ...

and

Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...

, and contains many familiar North American toad species such as the

American toad The American toad (''Anaxyrus americanus'') is a common species of toad found throughout Canada and the eastern United States. It is divided into three subspecies: the eastern American toad (''A. a. americanus''), the dwarf American toad (''A. a ...

Woodhouse's toad Woodhouse's toad (''Anaxyrus woodhousii'') is a medium-sized () true toad native to the United States and Mexico. There are three recognized subspecies. ''A. woodhousii'' tends to hybridize with '' Anaxyrus americanus'' where their ranges overla ...

, and the

western toad The western toad (''Anaxyrus boreas'') is a large toad species, between long, native to western North America. ''A. boreas'' is frequently encountered during the wet season on roads, or near water at other times. It can jump a considerable dista ...

. Most species in this genus were initially classified in ''

Bufo ''Bufo'' is a genus of true toads in the amphibian family Bufonidae. As traditionally defined, it was a wastebasket genus containing a large number of toads from much of the world but following taxonomic reviews most of these have been moved to ...

'', but were split due to their genetic divergence and geographic separation. Some authorities still consider ''Anaxyrus'' to be a

subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...

within ''Bufo''. However, other authorities have disputed this classification, as doing so would also require all morphologically distinct Old World toad species to also be placed in ''Bufo''.

Species

The following fossil taxa are also known, all of which were also previously placed in ''Bufo'': * †''

Anaxyrus defensor ''Anaxyrus'', containing the North American toads, is a genus of true toads in the family Bufonidae. The genus is endemic to North and Central America, and contains many familiar North American toad species such as the American toad, Woodhouse's ...

'' (Meylan, 2005) (Pliocene/early Pleistocene of

Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...

) * †''

Anaxyrus hibbardi ''Anaxyrus'', containing the North American toads, is a genus of true toads in the family Bufonidae. The genus is endemic to North and Central America, and contains many familiar North American toad species such as the American toad, Woodhouse's ...

'' (Taylor, 1937) (Late Miocene of

Kansas Kansas ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Nebraska to the north; Missouri to the east; Oklahoma to the south; and Colorado to the west. Kansas is named a ...

) * †''

Anaxyrus pliocompactilis ''Anaxyrus'', containing the North American toads, is a genus of true toads in the family Bufonidae. The genus is endemic to North and Central America, and contains many familiar North American toad species such as the American toad, Woodhouse's ...

'' (Wilson, 1968) (Late Miocene of Kansas, possibly synonymous with ''

Anaxyrus compactilis ''Anaxyrus compactilis'' (common name: plateau toad) is a species of toad in the family Bufonidae. It is endemic to Mexico. It is a little known species associated with desert and shrubland. It is threatened by habitat loss Habitat destruct ...

'' or '' Anaxyrus speciosus'') * †''

Anaxyrus repentinus ''Anaxyrus'', containing the North American toads, is a genus of true toads in the family Bufonidae. The genus is endemic to North and Central America, and contains many familiar North American toad species such as the American toad, Woodhouse's ...

'' (Tihen, 1962) (Mid-Late Pleistocene (

Rancholabrean The Rancholabrean North American Land Mammal Age on the geologic timescale is a North American faunal stage in the North American Land Mammal Ages chronology (NALMA),Sanders, A.E., R.E. Weems, and L.B. Albright III (2009) Formalization of the mid- ...

) of Kansas, possibly synonymous with ''

Anaxyrus woodhousii Woodhouse's toad (''Anaxyrus woodhousii'') is a medium-sized () true toad native to the United States and Mexico. There are three recognized subspecies. ''A. woodhousii'' tends to hybridize with ''Anaxyrus americanus'' where their ranges overlap ...

'' or '' Anaxyrus cognatus'') * †''

Anaxyrus rexroadensis ''Anaxyrus'', containing the North American toads, is a genus of true toads in the family Bufonidae. The genus is endemic to North and Central America, and contains many familiar North American toad species such as the American toad, Woodhouse's ...

'' (Tihen, 1962) (Pliocene/early Pleistocene of Kansas) * †''

Anaxyrus spongifrons ''Anaxyrus'', containing the North American toads, is a genus of true toads in the family Bufonidae. The genus is endemic to North and Central America, and contains many familiar North American toad species such as the American toad, Woodhouse's ...

'' (Tihen, 1962) (Late Miocene of Kansas) * †'' Anaxyrus suspectus'' (Tihen, 1962) (Pliocene/early Pleistocene (

Blancan The Blancan North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 4,750,000 to 1,806,000 years BP, a period of .Anaxyrus tiheni'' ( Auffenberg, 1957) (Late Miocene of Florida) * †''

Anaxyrus valentinensis ''Anaxyrus'', containing the North American toads, is a genus of true toads in the family Bufonidae. The genus is endemic to North and Central America, and contains many familiar North American toad species such as the American toad, Woodhouse's ...

'' (

Estes The surname "Estes" originates from the Middle English word est, meaning 'east'.https://forebears.io/surnames/estes The name "Estes" may also have its roots in ancient Greece, where it comes from the Greek "esthesis", which means "sensation" or "pe ...

& Tihen, 1964) (Middle Miocene (

Barstovian The Barstovian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 16,300,000 to 13,600,000 years BP, a period of . It is usua ...

) of

Nebraska Nebraska ( ) is a landlocked U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders South Dakota to the north; Iowa to the east and Missouri to the southeast, both across the Missouri River; Ka ...

)

References

External links

* * *

Species

References

External links

Further reading