Amphiprion Pacificus
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''Amphiprion pacificus'', is a species of
anemonefish Clownfishes or anemonefishes (genus ''Amphiprion'') are saltwater fishes found in the warm and tropical waters of the Indo-Pacific. They inhabit mainly coral reefs and have a distinctive colouration typically consisting of white vertical bars on ...
that is found in the western
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five Borders of the oceans, oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean, or, depending on the definition, to Antarctica in the south, and is ...
. Like all anemonefishes it forms a
symbiotic Symbiosis (Ancient Greek : living with, companionship < : together; and ''bíōsis'': living) is any type of a close and long-term biolo ...
mutualism with
sea anemone Sea anemones ( ) are a group of predation, predatory marine invertebrates constituting the order (biology), order Actiniaria. Because of their colourful appearance, they are named after the ''Anemone'', a terrestrial flowering plant. Sea anemone ...
s and is unaffected by the stinging tentacles of the host anemone. It is a sequential hermaphrodite with a strict sized based
dominance hierarchy In the zoological field of ethology, a dominance hierarchy (formerly and colloquially called a pecking order) is a type of social hierarchy that arises when members of animal social animal, social groups interact, creating a ranking system. Dif ...
: the female is largest, the breeding male is second largest, and the male non-breeders get progressively smaller as the hierarchy descends. They exhibit
protandry Sequential hermaphroditism (called dichogamy in botany) is one of the two types of hermaphroditism, the other type being simultaneous hermaphroditism. It occurs when the organism's sex changes at some point in its life. A sequential hermaphrodit ...
, meaning the breeding male will change to female if the sole breeding female dies, with the largest non-breeder becomes the breeding male. The fish's natural diet includes
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
.


Description

The body of ''A. pacificus'' is pinkish brown dark brown, usually grading to yellowish or orange on lower side of body including abdomen. It is a member of the skunk complex with the characteristic white stripe along the dorsal ridge line, from the midline of the snout through to the base of
caudal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only ...
. The fins are whitish to semi-translucent. It has 9 dorsal spines, 2 anal spines, 18-20 dorsal soft rays and 12-13 anal soft rays. The four specimens that were the basis of the description ranged from .


Color variations

Some anemonefish species have color variations based on geographic location, sex and host anemone. ''A. pacificus'', like other members of the skunk complex does not show any of these variations.


Similar species

'' A. akallopisos'' is almost identical, however there is a wide geographic separation such that geographic location is the easiest distinction. Despite the similarity in appearance, genetic results indicate that ''A. pacificus'' is well differentiated from ''A. akallopisos'' and is more closely related to '' A. sandaracinos'' which has a uniform coloration and the white stripe on the dorsal ridge extends onto the superior lip. 3 other species of anemonefish are found within its range, ''A. barberi'', ''A. chrysopterus'' and ''A. clarkii'', however these are easily distinguished by the absence of the white stripe on the dorsal ridge. Image:Amphiprion akallopisos 2010.JPG, ''A. akallopisos'' (Skunk anemonefish) Image:Amphiprion sandaracinos.jpg, ''A. sandaracinos'' (Orange anemonefish) showing the broader white stripe extending to the upper lip.


Distribution and Habitat

''A. pacificus'' is found the western Pacific at
Wallis Island Wallis () is a Polynesian atoll/island in the Pacific Ocean belonging to the French overseas collectivity (''collectivité d'outre-mer'', or ''COM'') of Wallis and Futuna. It lies north of Tonga, northeast of Fiji, east-northeast of the Hoorn ...
,
Tonga Tonga, officially the Kingdom of Tonga, is an island country in Polynesia, part of Oceania. The country has 171 islands, of which 45 are inhabited. Its total surface area is about , scattered over in the southern Pacific Ocean. accordin ...
,
Fiji Fiji, officially the Republic of Fiji, is an island country in Melanesia, part of Oceania in the South Pacific Ocean. It lies about north-northeast of New Zealand. Fiji consists of an archipelago of more than 330 islands—of which about ...
and
Samoan Islands The Samoan Islands () are an archipelago covering in the central Pacific Ocean, South Pacific, forming part of Polynesia and of the wider region of Oceania. Political geography, Administratively, the archipelago comprises all of the Samoa, Indep ...
, a regional hotspot of endemism. It is not common within its range and is generally found at depths of .


Host anemones

The relationship between anemonefish and their host sea anemones is not random and instead is highly
nested ''Nested'' is the seventh studio album by Bronx-born singer, songwriter, and pianist Laura Nyro. It was released in 1978 on Columbia Records. Following on from her extensive tour to promote 1976's ''Smile'', which resulted in the 1977 live albu ...
in structure. ''A. pacificus'' is highly specialised, being hosted by only 1 out of the 6 host anemones found in the region. ''A. pacificus'' is hosted by the following species of anemone: *''
Heteractis magnifica ''Radianthus magnifica'', also known by as magnificent sea anemone or Ritteri anemone, is a species of sea anemone in the family Stichodactylidae that is native to the Hawaii, USA, Indo-Pacific. Description The magnificent sea anemone is chara ...
'' magnificent sea anemone


Conservation status

Anemonefish and their host anemones are found on coral reefs and face similar
environmental issues Environmental issues are disruptions in the usual function of ecosystems. Further, these issues can be caused by humans (human impact on the environment) or they can be natural. These issues are considered serious when the ecosystem cannot recov ...
. Like
coral Corals are colonial marine invertebrates within the subphylum Anthozoa of the phylum Cnidaria. They typically form compact Colony (biology), colonies of many identical individual polyp (zoology), polyps. Coral species include the important Coral ...
s, anemone's contain intracellular
endosymbiont An endosymbiont or endobiont is an organism that lives within the body or cells of another organism. Typically the two organisms are in a mutualism (biology), mutualistic relationship. Examples are nitrogen-fixing bacteria (called rhizobia), whi ...
s,
zooxanthellae Zooxanthellae (; zooxanthella) is a colloquial term for single-celled photosynthetic organisms that are able to live in symbiosis with diverse marine invertebrates including corals, jellyfish, demosponges, and nudibranchs. Most known zooxanthell ...
, and can suffer from
bleaching Bleach is the generic name for any chemical product that is used industrially or domestically to remove color from (i.e. to whiten) fabric or fiber (in a process called bleaching) or to disinfect after cleaning. It often refers specifically t ...
due to triggers such as increased water temperature or
acidification Acidification may refer to: * Ocean acidification, decrease in the pH of the Earth's oceans * Freshwater acidification, atmospheric depositions and soil leaching of SOx and NOx * Soil acidification, buildup of hydrogen cations, which reduces the ...
. Characteristics known to elevate the risk of extinction are small geographic range, small local population and extreme habitat specialisation. ''A. pacificus'' has each of these characteristics, being an endemic species, with small local population and only one host anemone suggest that it is of particular conservation concern. This species was not evaluated in the 2012 release of the
IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...
.


References


External links

* * {{Taxonbar, from=Q219440 pacificus Fish described in 2010 Taxa named by Gerald R. Allen