Aeshna Petalura
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''Aeshna'', or the mosaic darners, is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
dragonflies A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of dragonflies are known. Most are tropical, with fewer species in temperate regions. Loss of wetland habitat threate ...
from the family
Aeshnidae Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and over 450 species. The family includes some of the largest dragonflies. Description Common worldwide or nearl ...
. Species within this genus are generally known as "hawkers" (
Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...
) or "darners" (
New World The term "New World" is used to describe the majority of lands of Earth's Western Hemisphere, particularly the Americas, and sometimes Oceania."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: ...
).


Description

These are relatively large dragonflies. Their thoraces and abdomens are brown in color, with blue or yellow stripes or spots on the thorax, and yellow, blue or green spots on the abdomen. Natalia von Ellenrieder's 2003 paper demonstrated that the
Holarctic The Holarctic realm is a biogeographic realm that comprises the majority of habitats found throughout the continents in the Northern Hemisphere. It corresponds to the floristic Boreal Kingdom. It includes both the Nearctic zoogeographical reg ...
and
Neotropic The Neotropical realm is one of the eight biogeographic realms constituting Earth's land surface. Physically, it includes the tropical terrestrial ecoregions of the Americas and the entire South American temperate zone. Definition In biogeogra ...
al species placed in this genus did not share a common ancestor, and proposed the latter be placed in the genus ''
Rhionaeschna ''Rhionaeschna'' is the scientific name of a genus of dragonflies from the family Aeshnidae Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and over 450 specie ...
''. The name ''Aeshna'' was coined by the Danish entomologist
Fabricius Fabricius (, ) is a surname. Notable people with the surname include: *people from the Ancient Roman Fabricia gens, gens Fabricia: **Gaius Fabricius Luscinus, the first of the Fabricii to move to Rome * Johann Goldsmid (1587–1616), known by his ...
in the 18th century. The name may have resulted from a printer's error in spelling the Greek ''Aechma'', "a spear". The spelling ''Aeschna'' has been intermittently used over a period of time, but is now abandoned for the original name ''Aeshna''. However, derived genus names (such as ''
Rhionaeschna ''Rhionaeschna'' is the scientific name of a genus of dragonflies from the family Aeshnidae Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and over 450 specie ...
'') retain the 'sch' spelling, as this is how they were first cited.


Species

Many species formerly included in ''Aeshna'' have been split into other genera, including ''
Afroaeschna ''Afroaeschna scotias'' is a species of dragonfly in the family Aeshnidae. It is found in Cameroon, Nigeria, Uganda, and Zambia. Its natural habitats are subtropical or tropical moist lowland forests and river A river is a natural strea ...
'', '' Andaeschna'', ''
Pinheyschna ''Pinheyschna'' is the scientific name of a genus of dragonflies from the family Aeshnidae. These relatively large dragonflies are also known as hawkers. Species The genus ''Pinheyschna'' includes the following species: *'' Pinheyschna meruensis ...
'', ''
Rhionaeschna ''Rhionaeschna'' is the scientific name of a genus of dragonflies from the family Aeshnidae Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and over 450 specie ...
'', and '' Zosteraeschna''. The genus ''Aeshna'' includes these species: *''
Aeshna affinis ''Aeshna affinis'', the southern migrant hawker or blue-eyed hawker, is a dragonfly found in southern Europe and Asia. It is in the family Aeshnidae and is very similar in appearance to '' A. mixta''. Identification ''Aeshna affinis'' is a sma ...
'' – southern migrant hawker, blue-eyed hawker *''
Aeshna athalia ''Aeshna athalia'' is a species of dragonfly in the genus ''Aeshna''. It is found in China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion ...
'' *''
Aeshna caerulea The azure hawker (''Aeshna caerulea'') is one of the smaller species of hawker dragonflies, (family Aeshnidae). The flight period is from late May to August It is about 62 mm long. Both sexes have azure blue spots on each abdominal segmen ...
'' – azure hawker *''
Aeshna canadensis ''Aeshna canadensis'', the Canada darner, is a species of dragonfly in the family Aeshnidae. It is common throughout southern Canada and the northern United States.Mead, Kurt. (2017). ''Dragonflies of the North Woods.'' 3rd edition. Duluth, MN: K ...
'' – Canada darner *''
Aeshna clepsydra ''Aeshna clepsydra'', the mottled darner, is a species of darner in the dragonfly family Aeshnidae Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and over 450 ...
'' – mottled darner *''
Aeshna constricta ''Aeshna constricta'', the lance-tipped darner, is a species of darner in the dragonfly family Aeshnidae Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and ov ...
'' – lance-tipped darner *''
Aeshna crenata ''Aeshna crenata'', the Siberian hawker, is a species of dragonfly in the family Aeshnidae. It is found in Belarus, Finland, Latvia, Lithuania, and Russia Russia, or the Russian Federation, is a country spanning Eastern Europe and North A ...
'' – Siberian hawker *''
Aeshna cyanea The southern hawker or blue hawker (''Aeshna cyanea'') is a species of hawker dragonfly. Distribution The species is one of the most common and most widespread dragonflies in Europe. The range in the Western Palearctic covers a large part of Eu ...
'' – blue hawker, southern hawker *''
Aeshna eremita ''Aeshna eremita'', the lake darner, is a species of dragonfly in the family Aeshnidae. It is found in Alaska, the Northern United States and across Canada Canada is a country in North America. Its Provinces and territories of Canada, ...
'' – lake darner *''
Aeshna frontalis ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' *''
Aeshna grandis The brown hawker (''Aeshna grandis'') is a species of large Palearctic dragonfly belonging to the family Aeshnidae, the hawkers. This species is found in Europe and temperate Asia, except the far east of Asia. Taxonomy The brown hawker was first ...
'' – brown hawker *''
Aeshna interrupta The variable darner (''Aeshna interrupta'') is a dragonfly of the family Aeshnidae, native from Alaska through the Northwest Territories to Newfoundland, south to New Hampshire and Michigan in the eastern United States, and to the mountains of New ...
'' – variable darner *''
Aeshna isoceles ''Aeshna isoceles'' (or ''isosceles'') is a small hawker dragonfly that is found in Europe, mostly around the Mediterranean, and the lowlands of North Africa. Its common name in English is green-eyed hawker. In the United Kingdom it is a localise ...
'' – Norfolk Hawker *''
Aeshna juncea The common hawker, moorland hawker or sedge darner (''Aeshna juncea'') is one of the larger species of hawker dragonflies. It is native to Palearctic (from Ireland to Japan) and northern North America. The flight period is from June to early Octo ...
'' – common hawker, sedge darner, moorland hawker *''
Aeshna mixta The migrant hawker (''Aeshna mixta'') is one of the smaller species of hawker dragonflies. It can be found away from water but for breeding it prefers still or slow-flowing water and can tolerate brackish sites. The flight period is from July to ...
'' – migrant hawker *''
Aeshna palmata The paddle-tailed darner (''Aeshna palmata'') is a species of dragonfly in the family Aeshnidae. It is common throughout western Canada and United States. This species is named after its distinctive paddle-shaped appendages. It lives in many habi ...
'' – paddle-tailed darner *''
Aeshna persephone ''Aeshna persephone'', Persephone's darner, is a species of dragonfly in the family Aeshnidae. It is found in northern Mexico and the southwestern United States. Its natural habitats are river A river is a natural stream of fresh wate ...
'' – Persephone's darner *'' Aeshna petalura'' *''
Aeshna septentrionalis ''Aeshna septentrionalis'', the azure darner, is a species of darner in the dragonfly family Aeshnidae Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and ove ...
'' – azure darner *''
Aeshna serrata The Baltic hawker (''Aeshna serrata'') is a species of hawker dragonfly A dragonfly is a flying insect belonging to the infraorder Anisoptera below the order Odonata. About 3,000 extant species of dragonflies are known. Most are tropica ...
'' – Baltic hawker *''
Aeshna shennong ''Aeshna'', or the mosaic darners, is a genus of dragonfly, dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large drag ...
'' *'' Aeshna sitchensis'' – zigzag darner *''
Aeshna soneharai ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' *''
Aeshna subarctica ''Aeshna subarctica'', the subarctic darner or bog hawker, is a species of dragonfly belonging to the family Aeshnidae, the darners and hawkers. This holarctic dragonfly is found in Europe and Northern Asia (excluding China) and North America. T ...
'' – bog hawker, subarctic darner *''
Aeshna tuberculifera ''Aeshna tuberculifera'', the black-tipped darner, is a species of darner in the dragonfly family Aeshnidae. It is found in North America. The IUCN conservation status of ''Aeshna tuberculifera'' is "LC", least concern, with no immediate threat ...
'' – black-tipped darner *''
Aeshna umbrosa The shadow darner (''Aeshna umbrosa'') is a species of dragonfly in the family Aeshnidae. It is found in almost all of Canada and most states in the United States. Identification The shadow darner is a large dragonfly with a length of . The ba ...
'' – shadow darner *'' Aeshna vercanica'' *'' Aeshna verticalis'' – green-striped darner *''
Aeshna viridis ''Aeshna viridis'', the green hawker, is a species of dragonfly in the family Aeshnidae. It is found in Austria, the Czech Republic, Denmark, Estonia, Finland, Germany, Hungary, Latvia, Lithuania, the Netherlands, Poland, Russia, Serbia and Mont ...
'' – green hawker *''
Aeshna walkeri ''Aeshna walkeri'', or Walker's darner, is a species of darner in the dragonfly family Aeshnidae Aeshnidae, also called aeshnids, hawkers, or darners, is a family of dragonflies, found nearly worldwide, with more than 50 genera and over 450 s ...
'' – Walker's darner *''
Aeshna williamsoniana ''Aeshna williamsoniana'', Williamson's darner, is a species of dragonfly in the family Aeshnidae. It is found in Belize, Costa Rica, Guatemala, Mexico, Panama, and possibly Honduras. Its natural habitats are subtropical or tropical moist monta ...
'' – Williamson's darner


Fossil species

Note that many fossil species in the genus were named at a time when many extant species now included in other genera were included in ''Aeshna''. Recent sources are not available to indicate whether these species belong in the new circumscription of ''Aeshna'' as used in this article. Fossil species include: *†''
Aeshna andancensis ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' – Late Miocene (
Turolian The Turolian age is a period of geologic time (9.0–5.3 Ma) within the Miocene used more specifically with European Land Mammal Ages. It precedes the Ruscinian age and follows the Vallesian age. The Turolian overlaps the Tortonian and Messinia ...
), France *†''
Aeshna cerdanica ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' – Late Miocene (
Vallesian The Vallesian age is a period of geologic time (11.6–9.0 Ma) within the Miocene used more specifically with European Land Mammal Ages. It precedes the Turolian age and follows the Astaracian age. The so-called Vallesian Crisis resulted in th ...
), Spain *†''
Aeshna dido ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' (Aeshnidae ''incertae sedis'') – Late Oligocene,
Rott Formation The Rott Formation is a geologic formation in Germany. It preserves fossils dating back to the Oligocene. See also * List of fossiliferous stratigraphic units in Germany See also * Lists of fossiliferous stratigraphic units in Europe * ...
, Germany *†''
Aeshna ghiandonii ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' – Late Miocene (
Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the fir ...
), Gessoso Formation, Italy *†''
Aeshna heterofasciata ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' (Anisoptera ''incertae sedis'') – Early Oligocene,
Salt Formation In common usage, salt is a mineral composed primarily of sodium chloride (NaCl). When used in food, especially in granulated form, it is more formally called table salt. In the form of a natural crystalline mineral, salt is also known as ro ...
, Germany *†''
Aeshna larvata ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' (Anisoptera ''incertae sedis'') – Oligocene,
Florissant Formation The Florissant Formation is a sedimentary geologic formation outcropping around Florissant, Teller County, Colorado. The formation is noted for the abundant and exceptionally preserved insect and plant fossils that are found in the mudstones an ...
, Colorado *†''
Aeshna messiniana ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' – Late Miocene (
Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the fir ...
), Gessoso Formation, Italy *†''
Aeshna multicellulata ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' – Late Miocene (
Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the fir ...
), Gessoso Formation, Italy *†''
Aeshna oligocenica ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' – terminal Oligocene,
Niveau du gypse d'Aix Formation The Niveau du gypse d'Aix Formation is a geologic formation in France. It preserves fossils dating back to the Paleogene period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly re ...
, France *†''
Aeshna ollivieri ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' – Late Oligocene, Campagne-Calavon Formation, France *†''
Aeshna paleocyanea ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' (Aeshnidae ''incertae sedis'') – Early Oligocene (Stampian), France *†''
Aeshna solida ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' – Oligocene,
Florissant Formation The Florissant Formation is a sedimentary geologic formation outcropping around Florissant, Teller County, Colorado. The formation is noted for the abundant and exceptionally preserved insect and plant fossils that are found in the mudstones an ...
, Colorado *†''
Aeshna theobaldi ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' (Aeshnidae ''incertae sedis'') – Early Pleistocene (
Villafranchian Villafranchian age ( ) is a period of geologic time (3.5–1.0 Ma) spanning the Late Pliocene and Early Pleistocene used more specifically with European Land Mammal Ages. Named by Italian geologist Lorenzo Pareto for a sequence of terrestrial se ...
), France *†''
Aeshna tyche ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' (Aeshnidae ''incertae sedis'') – Late Miocene (Sarmatian),
Upper Freshwater-Molasse Formation Upper may refer to: * Shoe upper or ''vamp'', the part of a shoe on the top of the foot * Stimulant, drugs which induce temporary improvements in either mental or physical function or both * ''Upper'', the original film title for the 2013 found fo ...
, Germany *†''
Aeshna voesendorfensis ''Aeshna'', or the mosaic darners, is a genus of dragonflies from the family Aeshnidae. Species within this genus are generally known as "hawkers" (Old World) or "darners" (New World). Description These are relatively large dragonflies. Th ...
'' – Late Miocene (
Messinian The Messinian is in the geologic timescale the last age or uppermost stage of the Miocene. It spans the time between 7.246 ± 0.005 Ma and 5.333 ± 0.005 Ma (million years ago). It follows the Tortonian and is followed by the Zanclean, the fir ...
), Austria


References


External links


''Aeshna''
BugGuide *Corbet, P. S. 1999. Dragonflies: Behavior and Ecology of Odonata. Cornell University Press, Ithaca, New York, USA, 829pp. *von Ellenrieder, N., 2003. A synopsis of the Neotropical species of'' 'Aeshna' ''Fabricius: the genus ''Rhionaeschna'' Förster (Odonata: Aeshnidae). - ''Tijdschrift voor Entomologie'' 146 (1): 67-207. {{Authority control Aeshnidae Taxa named by Johan Christian Fabricius Anisoptera genera