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Variable Checkerspot
The variable checkerspot or Chalcedon checkerspot (''Euphydryas chalcedona'') is a butterfly in the family Nymphalidae. It is found in western North America, where its range stretches from Alaska in the north to Baja California in the south and extends east through the Rocky Mountains into Colorado, Montana, New Mexico and Wyoming. The butterfly is usually brown or black with extensive white and yellow checkering and some red coloration on the dorsal wing. Adult wingspan is . Adult butterflies feed on nectar from flowers while larvae feed on a variety of plants including snowberry (''Symphoricarpos''), paintbrush (''Castilleja''), ''Buddleja'', '' Diplacus aurantiacus'' and ''Scrophularia californica''. During the breeding period, males congregate around larval host plants to encounter females. Males court female butterflies via physical displays.Rutowski, Ronald L., and George W. Gilchrist. "Courtship, Copulation and Oviposition in the Chalcedon Checkerspot,(Lepidoptera: Nymphali ...
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William O
William is a male given name of Germanic origin.Hanks, Hardcastle and Hodges, ''Oxford Dictionary of First Names'', Oxford University Press, 2nd edition, , p. 276. It became very popular in the English language after the Norman conquest of England in 1066,All Things William"Meaning & Origin of the Name"/ref> and remained so throughout the Middle Ages and into the modern era. It is sometimes abbreviated "Wm." Shortened familiar versions in English include Will, Wills, Willy, Willie, Bill, and Billy. A common Irish form is Liam. Scottish diminutives include Wull, Willie or Wullie (as in Oor Wullie or the play ''Douglas''). Female forms are Willa, Willemina, Wilma and Wilhelmina. Etymology William is related to the given name ''Wilhelm'' (cf. Proto-Germanic ᚹᛁᛚᛃᚨᚺᛖᛚᛗᚨᛉ, ''*Wiljahelmaz'' > German '' Wilhelm'' and Old Norse ᚢᛁᛚᛋᛅᚼᛅᛚᛘᛅᛋ, ''Vilhjálmr''). By regular sound changes, the native, inherited English form of the name shou ...
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Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism for the union of sperm with eggs. Flowers may facilitate outcrossing (fusion of sperm and eggs from different individuals in a population) resulting from cross-pollination or allow selfing (fusion of sperm and egg from the same flower) when self-pollination occurs. There are two types of pollination: self-pollination and cross-pollination. Self-pollination occurs when the pollen from the anther is deposited on the stigma of the same flower, or another flower on the same plant. Cross-pollination is when pollen is transferred from the anther of one flower to the stigma of another flower on a different individual of the same species. Self-pollination happens in flowers where the stamen and carpel mature at the same time, and are posi ...
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Order (biology)
Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. For some groups of organisms, their orders may fol ...
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Catalpol
Catalpol is an iridoid glucoside. This natural product falls in the class of iridoid glycosides, which are simply monoterpenes with a glucose molecule attached. Natural occurrence First isolated in 1962, catalpol was named for plants in the genus ''Catalpa'' in which it was discovered. Later in 1969, catalpol was found to be present in larger quantities in several plants in genus '' Rehmannia'' (Orobanchaceae). It is also found in plants belonging to several families, including, but not limited to, Scrophulariaceae, Lamiaceae (including scullcap), Plantaginaceae (''Plantago sp'') and Bignoniaceae, all of which being in the order Lamiales. Because they feed on these plants, variable checkerspot butterflies (''Euphydryas chalcedona'') contain high amounts of catalpol, which makes them unpalatable to predators and thus serves as a defense mechanism. Biosynthetic pathway Though first isolated in the 1960s, there has been very little investigation of the biosynthetic pathway ...
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Iridoid
Iridoids are a type of monoterpenoids in the general form of cyclopentanopyran, found in a wide variety of plants and some animals. They are biosynthetically derived from 8-oxogeranial. Iridoids are typically found in plants as glycosides, most often bound to glucose. The chemical structure is exemplified by iridomyrmecin, a defensive chemical produced by the ant genus ''Iridomyrmex'', for which iridoids are named. Structurally, they are bicyclic ''cis''-fused cyclopentane-pyrans. Cleavage of a bond in the cyclopentane ring gives rise to a subclass known as ''secoiridoids'', such as oleuropein and amarogentin. Occurrence The iridoids produced by plants act primarily as a defense against herbivores or against infection by microorganisms. The variable checkerspot butterfly also contains iridoids obtained through its diet which act as a defense against avian predators. To humans and other mammals, iridoids are often characterized by a deterrent bitter taste. Aucubin and catal ...
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Pupation
A pupa ( la, pupa, "doll"; plural: ''pupae'') is the life stage of some insects undergoing transformation between immature and mature stages. Insects that go through a pupal stage are holometabolous: they go through four distinct stages in their life cycle, the stages thereof being egg, larva, pupa, and imago. The processes of entering and completing the pupal stage are controlled by the insect's hormones, especially juvenile hormone, prothoracicotropic hormone, and ecdysone. The act of becoming a pupa is called pupation, and the act of emerging from the pupal case is called eclosion or emergence. The pupae of different groups of insects have different names such as #Chrysalis, ''chrysalis'' for the pupae of butterflies and ''tumbler'' for those of the mosquito family. Pupae may further be enclosed in other structures such as #Cocoon, cocoons, nests, or Animal shell, shells. Position in life cycle The pupal stage follows the larval stage and precedes adulthood (''imago'') in ...
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Diapause
In animal dormancy, diapause is the delay in development in response to regular and recurring periods of adverse environmental conditions.Tauber, M.J., Tauber, C.A., Masaki, S. (1986) ''Seasonal Adaptations of Insects''. Oxford University Press It is a physiological state with very specific initiating and inhibiting conditions. The mechanism is a means of surviving predictable, unfavorable environmental conditions, such as temperature extremes, drought, or reduced food availability. Diapause is observed in all the life stages of arthropods, especially insects. Embryonic diapause, a somewhat similar phenomenon, occurs in over 130 species of mammals, possibly even in humans, and in the embryos of many of the oviparous species of fish in the order Cyprinodontiformes. Activity levels of diapausing stages can vary considerably among species. Diapause may occur in a completely immobile stage, such as the pupae and eggs, or it may occur in very active stages that undergo extensive mig ...
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Mating Plug
A mating plug, also known as a copulation plug, sperm plug, vaginal plug, or sphragis (Latin, from Greek σφραγίς ''sphragis'', "a seal"), is gelatinous secretion used in the mating of some species. It is deposited by a male into a female genital tract, such as the vagina, and later hardens into a plug or glues the tract together. While females can expel the plugs afterwards, the male's sperm still gets a time advantage in getting to the egg, which is often the deciding factor in fertilization. The mating plug plays an important role in sperm competition and may serve as an alternative and more advantageous strategy to active mate guarding. In some species, such a passive mate-guarding strategy may reduce selection on large male size. Such a strategy may be advantageous because it would allow a male to increase reproductive success by spending more time pursuing new female mates rather than active mate guarding. Composition The mating plug of the '' Bombus terrestris'' was ...
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Spermatophore
A spermatophore or sperm ampulla is a capsule or mass containing spermatozoa created by males of various animal species, especially salamanders and arthropods, and transferred in entirety to the female's ovipore during reproduction. Spermatophores may additionally contain nourishment for the female, in which case it is called a nuptial gift, as in the instance of bush crickets. In the case of the toxic moth '' Utetheisa ornatrix'', the spermatophore includes sperm, nutrients, and pyrrolizidine alkaloids which prevent predation because it is poisonous to most organisms. However, in some species such as the Edith's checkerspot butterfly, the "gift" provides little nutrient value. The weight of the spermatophore transferred at mating has little effect on female reproductive output. Arthropods Spermatophores are the norm in arachnids and several soil arthropods. In various insects, such as bush crickets, the spermatophore is often surrounded by a proteinaceous spermatophyla ...
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Scrophularia Californica
''Scrophularia californica'' is a flowering plant in the figwort family which is known by the common names California figwort and California bee plant. It is native to the western United States, including many habitats in California, and in British Columbia. Description ''Scrophularia californica'' is an unassuming plant with triangular, toothed, blue-green leaves in pairs opposite each other on a spindly, squared stem. The brownish-magenta flowers are rounded, hollow buds about a centimeter long with two long upper lobes. This species is a strong bee attractant and also serves as a host plant In biology and medicine, a host is a larger organism that harbours a smaller organism; whether a parasitic, a mutualistic, or a commensalist ''guest'' ( symbiont). The guest is typically provided with nourishment and shelter. Examples include ... for variable checkerspot larvae.Williams, K. S., D. E. Lincoln, and P. R. Ehrlich. "The Coevolution of Euphydryas Chalcedona Butterfli ...
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Diplacus Aurantiacus
''Diplacus aurantiacus'', the sticky monkey-flower or orange bush monkey-flower, is a flowering plant that grows in a subshrub form, native to southwestern North America from southwestern Oregon south through most of California. It is a member of the lopseed family, Phrymaceae. It was formerly known as ''Mimulus aurantiacus''.''Mimulus aurantiacus''.
The Jepson Manual.


Description

''Diplacus aurantiacus'' grows up to 1.2 meters (4 feet) tall, has deep green, sticky leaves 3 to 7 centimeters long and up to a centimeter broad and flowering stems that grow vertically. The

Buddleja
''Buddleja'' (; ''Buddleia''; also historically given as ''Buddlea'') is a genus comprising over 140 species of flowering plants endemic to Asia, Africa, and the Americas. The generic name bestowed by Linnaeus posthumously honoured the Reverend Adam Buddle (1662–1715), an English botanist and rector, at the suggestion of Dr. William Houstoun. Houstoun sent the first plants to become known to science as buddleja ( ''B. americana'') to England from the Caribbean about 15 years after Buddle's death. Nomenclature The botanic name has been the source of some confusion. By modern practice of botanical Latin, the spelling of a generic name made from 'Buddle' would be ''Buddleia'', but Linnaeus in his '' Species Plantarum'' of 1753 and 1754 spelled it ''Buddleja'', with the long i between two vowels, common in early modern orthography.Linnaei, C. (1753). ''Species plantarum''. Impensis Laurentii Salvii, Stockholm. The pronunciation of the long i in ''Buddleja'' as ''j'' is a co ...
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