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Trechnotheria
Trechnotheria is a group of mammals that includes the therians and some fossil mammals from the Mesozoic Era. It includes both the extinct symmetrodonts and the living Cladotheria. Trechnotheria has been assigned various ranks, but was originally described as a "superlegion" by the naming authority. A later node-based definition for Trechnotheria was the clade comprising the last common ancestor of '' Zhangheotherium'' and living therian mammals, and all its descendants. Alternatively it can be defined as the group with true symmetrodontia, being defined as an apomorphy rather than being defined as a node, the redefinition includes the Tinodontids. Characteristics Like most Mesozoic mammal groups, early trechnotherians are known mainly from their teeth. Hence, one of the most prominent features of this group is the "hypertrophied postvallum/prevallid shearing mechanism", along with other dental characters. Features of the shoulder blade, tibia, humerus, and ankle joint a ...
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Spalacotheriida
Symmetrodonta is a group of Mesozoic mammals and mammal-like synapsids characterized by the triangular aspect of the Molar (tooth), molars when viewed from above, and the absence of a well-developed talonid. The traditional group of 'symmetrodonts' ranges in age from the latest Triassic to the Late Cretaceous, but most research in the last 20-30 years has concluded that they are not a true taxonomic group, but include several unrelated branches of the mammal tree. Despite this, the name is still used informally by some researchers for convenience, usually restricted to the spalacotheriids and zhangheotheriids. Martin, T., 2018. 6. Mesozoic mammals—early mammalian diversity and ecomorphological adaptations. In Mammalian evolution, diversity and systematics (pp. 199-300). De Gruyter. There are some symmetrodonts with acutely-triangulated molar cusps (“acute-angled symmetrodonts”) that seem to form a true monophyletic group, and lasted from the Early Cretaceous to the Campanian ...
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Zhangheotheriidae
Zhangheotheriidae is a possibly paraphyletic family of "symmetrodont" mammals that is currently known from Early Cretaceous deposits in China and Russia. Six genera are currently recognized, ''Anebodon'', ''Kiyatherium'', ''Maotherium'', ''Ningchengodon'', ''Origolestes'', and ''Zhangheotherium''. References

Symmetrodonta Prehistoric mammal families Early Cretaceous mammals of Asia {{cretaceous-mammal-stub ...
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Zatheria
Cladotheria is a clade (sometimes ranked as a Legion (taxonomy), legion) of mammals. It contains modern therian mammals (marsupials and Placentalia, placentals) and several extinct groups, such as the "dryolestoids", amphitheriids and peramurids. The clade was named in 1975 by Malcolm McKenna. In 2002, it was defined as a node-based taxon containing "the common ancestor of dryolestids and living therians, plus all its descendants". A different, stem-based taxon, stem-based definition was given in 2013, in which Cladotheria contains all taxa that are closer to ''Mus musculus'' (the house mouse) than to the "symmetrodont" ''Spalacotherium tricuspidens''. Cladotheria incorporates a set of nested mammal clades culminating in Tribosphenida (also known as Boreosphenida), mammals with fully Tribosphenic molar, tribosphenic teeth such as therians and a few of their closest relatives. The clade Prototribosphenida includes "the common ancestor of ''Vincelestes'' and living therians, plus al ...
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Symmetrodont
Symmetrodonta is a group of Mesozoic mammals and mammal-like synapsids characterized by the triangular aspect of the molars when viewed from above, and the absence of a well-developed talonid. The traditional group of 'symmetrodonts' ranges in age from the latest Triassic to the Late Cretaceous, but most research in the last 20-30 years has concluded that they are not a true taxonomic group, but include several unrelated branches of the mammal tree. Despite this, the name is still used informally by some researchers for convenience, usually restricted to the spalacotheriids and zhangheotheriids. Martin, T., 2018. 6. Mesozoic mammals—early mammalian diversity and ecomorphological adaptations. In Mammalian evolution, diversity and systematics (pp. 199-300). De Gruyter. There are some symmetrodonts with acutely-triangulated molar cusps (“acute-angled symmetrodonts”) that seem to form a true monophyletic group, and lasted from the Early Cretaceous to the Campanian, although ...
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Symmetrodonts
Symmetrodonta is a group of Mesozoic mammals and mammal-like synapsids characterized by the triangular aspect of the molars when viewed from above, and the absence of a well-developed talonid. The traditional group of 'symmetrodonts' ranges in age from the latest Triassic to the Late Cretaceous, but most research in the last 20-30 years has concluded that they are not a true taxonomic group, but include several unrelated branches of the mammal tree. Despite this, the name is still used informally by some researchers for convenience, usually restricted to the spalacotheriids and zhangheotheriids. Martin, T., 2018. 6. Mesozoic mammals—early mammalian diversity and ecomorphological adaptations. In Mammalian evolution, diversity and systematics (pp. 199-300). De Gruyter. There are some symmetrodonts with acutely-triangulated molar cusps (“acute-angled symmetrodonts”) that seem to form a true monophyletic group, and lasted from the Early Cretaceous to the Campanian, although Zha ...
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Boreosphenida
Tribosphenida is a clade of mammals that includes the ancestor of ''Hypomylos'', Aegialodontia and Theria (the last common ancestor of marsupials and placentals plus all of its descendants). It belongs to the group Zatheria. The current definition of Tribosphenida is more or less synonymous with Boreosphenida. Characteristics Tribosphenid mammals were originally grouped on the basis of triangular or V-shaped (Molar (tooth)#Tribosphenic, tribosphenic) molars. The relationship of the also tribosphenic australosphenidans, a group of mammals from the Jurassic-Cretaceous of the Southern Hemisphere often suggested to be close relatives of living Monotreme, monotremes, has been questioned, and it has been argued that they developed tribosphenic molars independently from those of "true" tribosphenidans. Some authors have alternatively continued to argue that non-monotreme australosphenidans are in fact true tribosphenidans unrelated to monotremes. "True" unambiguous members of Tribospheni ...
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Dryolestida
Dryolestida is an extinct order of mammals, known from the Jurassic and Cretaceous. They are considered basal members of the clade Cladotheria, close to the ancestry of therian mammals. It is also believed that they developed a fully mammalian jaw and also had the three middle ear bones. Most members of the group, as with most Mesozoic mammals, are only known from fragmentary tooth and jaw remains. The group contains Dryolestidae and the possibly paraphyletic Paurodontidae, and some other unplaced genera, which were small insectivores, known from the Middle Jurassic to Early Cretaceous of Laurasia, primarily Europe and North America, with a single record from Asia. During the Late Jurassic in North America and from the Late Jurassic to Early Cretaceous in Europe, they were among the most diverse mammal groups. They have sometimes been placed as part of a broader group, the "Dryolestoidea", which typically includes Dryolestida and Meridiolestida, a diverse group of mammals in ...
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Cladotheria
Cladotheria is a clade (sometimes ranked as a legion) of mammals. It contains modern therian mammals (marsupials and placentals) and several extinct groups, such as the "dryolestoids", amphitheriids and peramurids. The clade was named in 1975 by Malcolm McKenna. In 2002, it was defined as a node-based taxon containing "the common ancestor of dryolestids and living therians, plus all its descendants". A different, stem-based definition was given in 2013, in which Cladotheria contains all taxa that are closer to ''Mus musculus'' (the house mouse) than to the " symmetrodont" '' Spalacotherium tricuspidens''. Cladotheria incorporates a set of nested mammal clades culminating in Tribosphenida (also known as Boreosphenida), mammals with fully tribosphenic teeth such as therians and a few of their closest relatives. The clade Prototribosphenida includes "the common ancestor of '' Vincelestes'' and living therians, plus all of its descendants". Apart from tribosphenids, Prototrib ...
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Symmetrodonta
Symmetrodonta is a group of Mesozoic mammals and mammal-like synapsids characterized by the triangular aspect of the molars when viewed from above, and the absence of a well-developed talonid. The traditional group of 'symmetrodonts' ranges in age from the latest Triassic to the Late Cretaceous, but most research in the last 20-30 years has concluded that they are not a true taxonomic group, but include several unrelated branches of the mammal tree. Despite this, the name is still used informally by some researchers for convenience, usually restricted to the spalacotheriids and zhangheotheriids. Martin, T., 2018. 6. Mesozoic mammals—early mammalian diversity and ecomorphological adaptations. In Mammalian evolution, diversity and systematics (pp. 199-300). De Gruyter. There are some symmetrodonts with acutely-triangulated molar cusps (“acute-angled symmetrodonts”) that seem to form a true monophyletic group, and lasted from the Early Cretaceous to the Campanian, although ...
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Pappotheriidae
''Pappotherium'' is an extinct genus of mammals from the Albian (early Cretaceous) of Texas, US, known from a fossilized maxilla fragment bearing two tribosphenic molars, discovered within the Glen Rose Formation near Decatur, Wise County, Texas. The fossil was discovered by Bob H. Slaughter within some deposits dating back to 112.6 – 109 million years ago. On the basis of the morphology of the molars' cusps, in 1965 Slaughter established the new genus ''Pappotherium'' and the new species ''P. pattersoni''; he also created an apposite family, Pappotheriidae. Both this family and the genus are nowadays still monotypic. Slaughter argued that ''Pappotherium'' should have been a basal form close to the metatherian-eutherian divergence point; this mammal likely was an arboreal insectivore. Etymologically speaking, the name ''Pappotherium'' is a compound of the Latin words ''pappus'' (from ancient Greek πάππος, ''páppos'', “grandfather”) and ''therium'' (from ancient Gr ...
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Aegialodontia
Aegialodontia is a clade of extinct early mammals, close to the origin of Boreosphenida. The clade includes some of the oldest known tribosphenic taxa, until the discovery of '' Tribactonodon'' from the Berriasian Durlston Formation in 2001, '' Aegialodon'' from the Valanginian Wadhurst Clay Formation was the oldest taxon with the tooth form. The Aptian to Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ... taxon '' Kielantherium'' from Mongolia, formerly a synonym of ''Aegialodon'', is also within the group, sister to ''Aegialodon'' within Aegialodontidae. References Prehistoric mammals Tribosphenida {{Cretaceous-mammal-stub ...
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Theria
Theria ( or ; ) is a scientific classification, subclass of mammals amongst the Theriiformes. Theria includes the eutherians (including the Placentalia, placental mammals) and the metatherians (including the marsupials) but excludes the egg-laying monotremes and various extinct mammals evolving prior to the common ancestor of placentals and marsupials. Characteristics Therians give birth to live young without a shelled egg (biology), egg. This is possible thanks to key proteins called Syncytin-1, syncytins which allow exchanges between the mother and its offspring through a placenta, even Marsupial#Reproductive system, rudimental ones such as in marsupials. Genetic studies have suggested a viral origin of syncytins through the Endogenous retrovirus, endogenization process. The marsupials and the placentals evolved from a common therian ancestor that gave live birth by suppressing the mother's immune system. While the marsupials continued to give birth to an underdeveloped fe ...
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