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Phylogeny Of Malacostraca
Phylogeny of Malacostraca is the evolutionary relationships of the largest of the six classes of crustaceans, containing about 40,000 living species, divided among 16 orders. Its members display a great diversity of body forms. Although the class Malacostraca is united by a number of well-defined and documented features, which were recognised a century ago by William Thomas Calman in 1904, the phylogenetic relationship (the evolutionary tree) of the orders which compose this class is unclear due to the vast diversity present in their morphology. Molecular studies have attempted to infer the phylogeny of this clade, resulting in phylogenies which have a limited amount of morphological support. To resolve a well-supported eumalacostracan phylogeny and obtain a robust tree, it will be necessary to look beyond the most commonly utilized sources of data (nuclear ribosomal and mitochondrial sequences). Features The Malacostraca is assumed to be monophyletic due to several common morp ...
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Crustacean
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, amphipoda, amphipods and mantis shrimp. The crustacean group can be treated as a subphylum under the clade Mandibulata. It is now well accepted that the Hexapoda, hexapods emerged deep in the Crustacean group, with the completed group referred to as Pancrustacea. Some crustaceans (Remipedia, Cephalocarida, Branchiopoda) are more closely related to insects and the other hexapods than they are to certain other crustaceans. The 67,000 described species range in size from ''Stygotantulus, Stygotantulus stocki'' at , to the Japanese spider crab with a leg span of up to and a mass of . Like other arthropods, crustaceans have an exoskeleton, which they ecdysis, moult to grow. They are distinguished from other groups of arthropods, such as insec ...
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Phyllocarida
Phyllocarida is a subclass of crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...s, comprising the extant order Leptostraca and the extinct orders Hymenostraca and Archaeostraca. See also *'' Ceratiocaris'' *'' Cinerocaris'' *'' Vladicaris'' References External links Malacostraca Arthropod subclasses Paleozoic crustaceans Mesozoic crustaceans Cenozoic crustaceans Extant Cambrian first appearances Taxa described in 1879 Taxa named by Alpheus Spring Packard {{Malacostraca-stub ...
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Carapace
A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron. Crustaceans In crustaceans, the carapace functions as a protective cover over the cephalothorax (i.e., the fused head and thorax, as distinct from the abdomen behind). Where it projects forward beyond the eyes, this projection is called a rostrum. The carapace is calcified to varying degrees in different crustaceans. Zooplankton within the phylum Crustacea also have a carapace. These include Cladocera, ostracods, and isopods, but isopods only have a developed "cephalic shield" carapace covering the head. Arachnids In arachnids, the carapace is formed by the fusion of prosomal tergites into a single plate which carries the eyes, ocularium, ozopores (a pair of openings of the scent gland of Opiliones) a ...
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Uropod
Uropods are posterior appendages found on a wide variety of crustaceans. They typically have functions in locomotion. Definition Uropods are often defined as the appendages of the last body segment of a crustacean. An alternative definition suggested by Frederick R. Schram restricts the term to those structures arising from the segment before the anal segment (the segment which carries the anus). Under this latter definition, the appendages of the anal segment are caudal rami, which are analogous to uropods. Form Uropods are typically biramous – comprising an endopod and an exopod. The exopod is typically the larger, and may be divided in two by a transverse suture known as the diaeresis. The uropods may work in concert with the telson The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment o ...
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Telson
The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on account of not arising in the embryo from teloblast areas as other segments. It never carries any appendages, but a forked "tail" called the caudal furca may be present. The shape and composition of the telson differs between arthropod groups. Crustaceans In lobsters, shrimp and other decapods, the telson, along with the uropods, forms the tail fan. This is used as a paddle in the caridoid escape reaction ("lobstering"), whereby an alarmed animal rapidly flexes its tail, causing it to dart backwards. Krill can reach speeds of over 60 cm per second by this means. The trigger time to optical stimulus is, in spite of the low temperatures, only 55  milliseconds. In the Isopoda and Tanaidacea (superorder Peracarida), the last ...
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Pleon
The decapod (crustaceans such as a crab, lobster, shrimp or prawn) is made up of 20 body segments grouped into two main body parts: the cephalothorax and the pleon (abdomen). Each segment may possess one pair of appendages, although in various groups these may be reduced or missing. They are, from head to tail: Cephalothorax Head #antennules # antennae # mandibles # first maxillae # second maxillae The head also bears the (usually stalked) compound eyes. The distal portion of a mandible or maxilla which has a sensory function is known as a palp. Thorax / pereon #first maxillipeds #second maxillipeds #third maxillipeds #first pereiopods #second pereiopods #third pereiopods #fourth pereiopods #fifth pereiopods Maxillipeds are appendages modified to function as mouthparts. Particularly in the less advanced decapods, these can be very similar to the pereiopods. Pereiopods are primarily walking legs and are also used for gathering food. They are also the ten legs from which decapod ...
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Thorax
The thorax or chest is a part of the anatomy of humans, mammals, and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main divisions of the creature's body, each of which is in turn composed of multiple segments. The human thorax includes the thoracic cavity and the thoracic wall. It contains organs including the heart, lungs, and thymus gland, as well as muscles and various other internal structures. Many diseases may affect the chest, and one of the most common symptoms is chest pain. Etymology The word thorax comes from the Greek θώραξ ''thorax'' " breastplate, cuirass, corslet" via la, thorax. Plural: ''thoraces'' or ''thoraxes''. Human thorax Structure In humans and other hominids, the thorax is the chest region of the body between the neck and the abdomen, along with its internal organs and other contents. It is mostly protected and supported by the rib ...
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Cephalon (arthropod Anatomy)
The cephalon is the head section of an arthropod. It is a tagma, i.e., a specialized grouping of arthropod segments. The word cephalon derives from the Greek κεφαλή (kephalē), meaning "head". Insects In insects, ''head'' is a preferred term. The insect head consists of five segments, including three (the labial, maxillary and mandibular) necessary for food uptake, which are altogether known as the gnathocephalon and house the suboesophageal ganglion of the brain, as well as the antennal segment, and an ocular segment, as well as a non segmented fused section of the head where the archicerebrum is housed known as the acron. See also arthropod head problem. Chelicerates and crustaceans In chelicerates and crustaceans, the cephalothorax is derived from the fusion of the cephalon and the thorax, and is usually covered by a single unsegmented carapace. In relation with the arthropod head problem, phylogeny studies show that members of the Malacostraca class of crus ...
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Tagma (biology)
In biology, a tagma (Greek: τάγμα, plural tagmata – τάγματα - body of soldiers; battalion) is a specialized grouping of multiple segments or metameres into a coherently functional morphological unit. Familiar examples are the head, the thorax, and the abdomen of insects. The segments within a tagma may be either fused (such as in the head of an insect) or so jointed as to be independently moveable (such as in the abdomen of most insects). Usually the term is taken to refer to tagmata in the morphology of members of the phylum Arthropoda, but it applies equally validly in other phyla, such as the Chordata. In a given taxon the names assigned to particular tagmata are in some sense informal and arbitrary; for example, not all the tagmata of species within a given subphylum of the Arthropoda are homologous to those of species in other subphyla; for one thing they do not all comprise corresponding somites, and for another, not all the tagmata have closely analogou ...
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Syncarida
Syncarida is a superorder of crustaceans, comprising the two extant orders Anaspidacea and Bathynellacea, and the extinct order Palaeocaridacea. Fifty-nine living genera are known, in six families: ;Anaspidacea Calman, 1904 *Anaspididae Thomson, 1893 *Koonungidae Sayce, 1908 *Psammaspididae Schminke, 1974 *Stygocarididae Stygocarididae is a family of crustaceans belonging to the order Anaspidacea Anaspidacea is an order of crustaceans, comprising eleven genera in four families. Species in the family Anaspididae vary from being strict stygobionts (only living ... Noodt, 1963 ; Bathynellacea Chappuis, 1915 * Bathynellidae Grobben, 1904 * Parabathynellidae Noodt, 1965 References * Arthropod superorders Taxa named by Alpheus Spring Packard {{Malacostraca-stub ...
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Hoplocarida
Hoplocarida is a subclass of crustaceans. The only extant members are the mantis shrimp (Stomatopoda), but two other orders existed in the Palaeozoic: Aeschronectida Aeschronectida is an extinct order of mantis shrimp-like crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapods, seed shrimp, branchiopods, fish lice, krill, remipedes, isopods, ... and Palaeostomatopoda. References Malacostraca Arthropod subclasses {{Malacostraca-stub ...
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Peracarida
The superorder Peracarida is a large group of malacostracan crustaceans, having members in marine, freshwater, and terrestrial habitats. They are chiefly defined by the presence of a brood pouch, or ''marsupium'', formed from thin flattened plates (oostegites) borne on the basalmost segments of the legs. Peracarida is one of the largest crustacean taxa and includes about 12,000 species. Most members are less than in length, but the largest is probably the giant isopod (''Bathynomus giganteus'') which can reach . The earliest known perecaridian was '' Oxyuropoda ligioides'', a fossil of which has been found dating to the Late Devonian (more than 360 mya) of Ireland. Characteristics The most obvious characteristic of the group is the marsupium in females. This brood pouch is enclosed by the large, flexible oostergites, bristly flaps which extend from the basal segments of the thoracic appendages, which form the floor of a chamber roofed by the animal's sternum. This chamber is ...
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