Periostracum
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Periostracum
The periostracum ( ) is a thin, organic coating (or "skin") that is the outermost layer of the shell of many shelled animals, including molluscs and brachiopods. Among molluscs, it is primarily seen in snails and clams, i.e. in gastropods and bivalves, but it is also found in cephalopods such as ''Allonautilus scrobiculatus''. The periostracum is an integral part of the shell, and it forms as the shell forms, along with the other shell layers. The periostracum is used to protect the organism from corrosion. The periostracum is visible as the outer layer of the shell of many molluscan species from terrestrial, freshwater, and marine habitats, and may be seen in land snails, river mussels, and other kinds of freshwater bivalves, as well as in many kinds of marine shelled molluscs. The word ''periostracum'' means "around the shell", meaning that the periostracum is wrapped around what is usually the more calcareous part of the shell. Technically, the calcareous part of the ...
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Brachiopod
Brachiopods (), phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically-oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a ...
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Gastropod Shell
The gastropod shell is part of the body of a gastropod or snail, a kind of mollusc. The shell is an exoskeleton, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some gastropods appear shell-less (slugs) but may have a remnant within the mantle, or in some cases the shell is reduced such that the body cannot be retracted within it ( semi-slug). Some snails also possess an operculum that seals the opening of the shell, known as the aperture, which provides further protection. The study of mollusc shells is known as conchology. The biological study of gastropods, and other molluscs in general, is malacology. Shell morphology terms vary by species group. Shell layers The gastropod shell has three major layers secreted by the mantle. The calcareous central layer, tracum, is typically made of calcium carbonate precipitated into an organic matrix known as conchiolin. The outermost layer is the periostr ...
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Allonautilus Scrobiculatus
''Allonautilus scrobiculatus'', also known as the crusty nautilus or fuzzy nautilus, is a species of nautilus native to the waters around New Guinea, specifically New Britain and Milne Bay, and the Solomon Islands. ''A. scrobiculatus'' is recognizable by the large open umbilicus, which is around 20% of the shell diameter at its widest point. This species, along with the closely related '' A. perforatus'', were originally placed in the genus ''Nautilus'', but have recently been given their own genus on account of significant morphological differences.Ward, P.D. & W.B. Saunders 1997''Allonautilus'': a new genus of living nautiloid cephalopod and its bearing on phylogeny of the Nautilida ''Journal of Paleontology'' 71(6): 1054–1064. The most obvious are features of the shell, including crease and an encrusting layer ( periostracum) that covers most of the shell. Gills and reproductive structures also differ significantly from members of the genus ''Nautilus''. The shell is usual ...
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Bivalvia
Bivalvia (), in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of marine and freshwater molluscs that have laterally compressed bodies enclosed by a shell consisting of two hinged parts. As a group, bivalves have no head and they lack some usual molluscan organs, like the radula and the odontophore. They include the clams, oysters, cockles, mussels, scallops, and numerous other families that live in saltwater, as well as a number of families that live in freshwater. The majority are filter feeders. The gills have evolved into ctenidia, specialised organs for feeding and breathing. Most bivalves bury themselves in sediment, where they are relatively safe from predation. Others lie on the sea floor or attach themselves to rocks or other hard surfaces. Some bivalves, such as the scallops and file shells, can swim. The shipworms bore into wood, clay, or stone and live inside these substances. The shell of a bivalve is composed o ...
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Conchiolin
Conchiolins (sometimes referred to as conchins) are complex proteins which are secreted by a mollusc's outer epithelium (the mantle). These proteins are part of a matrix of organic macromolecules, mainly proteins and polysaccharides, that assembled together form the microenvironment where crystals nucleate and grow. This organic matrix also holds and binds to the crystals of aragonite which give such shells their stiffness. The ions necessary to form calcium carbonate are also secreted by the mantle, but it is the tailored environment created by the organic matrix which causes aragonite (rather than calcite) crystals to nucleate, in much the same way that collagen nucleates hydroxyapatite crystals. Conchiolin serves as a relatively flexible, crack-deflecting matrix for the mineral aggregate particles; its strength and the strong bonding of perlucin can in some cases (such as in the formation of nacre) give the finished material an impressive level of toughness. As ...
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Mollusc
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The gastr ...
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Trochulus
''Trochulus'' is a genus of small air-breathing land snails, terrestrial pulmonate gastropod mollusks in the subfamily subfamily Trochulininae of the family Hygromiidae, the hairy snails and their allies. Taxonomy ''Trichia'' Hartmann, 1840 is a junior synonym of ''Trochulus'' Chemnitz, 1786. Nearly every malacological work prior to 2006 used the name ''Trichia'' instead of the (now considered valid) name ''Trochulus''. The genus '' Plicuteria'' Schileyko, 1978 has been once recognized as a subgenus within ''Trochulus'' by Schileyko (1978). Based on molecular analyses, ''Trochulus lubomirski'' does not belong to the genus ''Trochulus''. ''Trochulus lubomirski'' (Ślósarski, 1881) is now recognized as '' Plicuteria lubomirski'' (Ślósarski, 1881). Species The speciation centre for the genus ''Trochulus'' is in the Alps. The type species of this genus is '' Trochulus hispidus''. Species within the genus ''Trochulus'' include: * '' Trochulus alpicola'' (Eder, 1921) ...
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Cuticle
A cuticle (), or cuticula, is any of a variety of tough but flexible, non-mineral outer coverings of an organism, or parts of an organism, that provide protection. Various types of "cuticle" are non- homologous, differing in their origin, structure, function, and chemical composition. Human anatomy In human anatomy, "cuticle" can refer to several structures, but it is used in general parlance, and even by medical professionals, to refer to the thickened layer of skin surrounding fingernails and toenails (the eponychium), and to refer to the superficial layer of overlapping cells covering the hair shaft ( cuticula pili), consisting of dead cells, that locks the hair into its follicle. It can also be used as a synonym for the epidermis, the outer layer of skin. Cuticle of invertebrates In zoology, the invertebrate cuticle or cuticula is a multi-layered structure outside the epidermis of many invertebrates, notably roundworms and arthropods, in which it forms an exoskeleto ...
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Varnish
Varnish is a clear transparent hard protective coating or film. It is not a stain. It usually has a yellowish shade from the manufacturing process and materials used, but it may also be pigmented as desired, and is sold commercially in various shades. Varnish is primarily used as a wood finish where, stained or not, the distinctive tones and grains in the wood are intended to be visible. Varnish finishes are naturally glossy, but satin/semi-gloss and flat sheens are available. History The word "varnish" comes from Mediaeval Latin ''vernix'', meaning odorous resin, itself derived from Middle Greek ''berōnikón'' or ''beroníkē'', meaning amber or amber-colored glass. A false etymology traces the word to the Greek '' Berenice'', the ancient name of modern Benghazi in Libya, where the first varnishes in the Mediterranean area were supposedly used and where resins from the trees of now-vanished forests were sold. Early varnishes were developed by mixing resin—pine sap, fo ...
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Cone Snail
A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines connecting a common point, the apex, to all of the points on a base that is in a plane that does not contain the apex. Depending on the author, the base may be restricted to be a circle, any one-dimensional quadratic form in the plane, any closed one-dimensional figure, or any of the above plus all the enclosed points. If the enclosed points are included in the base, the cone is a solid object; otherwise it is a two-dimensional object in three-dimensional space. In the case of a solid object, the boundary formed by these lines or partial lines is called the ''lateral surface''; if the lateral surface is unbounded, it is a conical surface. In the case of line segments, the cone does not extend beyond the base, while in the case of half-line ...
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Malacologist
Malacology is the branch of invertebrate zoology that deals with the study of the Mollusca (mollusks or molluscs), the second-largest phylum of animals in terms of described species after the arthropods. Mollusks include snails and slugs, clams, and cephalopods, along with numerous other kinds, many of which have shells. One division of malacology, conchology, is devoted to the study of mollusk shells. Malacology derives . Fields within malacological research include taxonomy, ecology and evolution. Applied malacology studies medical, veterinary, and agricultural applications; for example, mollusks as vectors of disease, as in schistosomiasis. Archaeology employs malacology to understand the evolution of the climate, the biota of the area, and the usage of the site. In 1681, Filippo Bonanni wrote the first book ever published that was solely about seashells, the shells of marine mollusks. The book was entitled: In 1868, the German Malacological Society was founded. Zoologica ...
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