Megalichthyid
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Megalichthyid
Megalichthyidae is an extinct family of tetrapodomorphs which lived from the Middle–Late Devonian to the Early Permian. They are known primarily from freshwater deposits, mostly in the Northern Hemisphere (Europe, the Middle East, North Africa and North America), but one genus (''Cladarosymblema'') is known from Australia, and the possible megalichthyid ''Mahalalepis'' is from Antarctica. Description Megalichthyids were fairly primitive tetrapodomorphs, retaining a largely fish-like appearance. Like some other primitive sarcopterygians, their bodies were covered in rhomboid scales that possessed a layer of cosmine (a porous, mineralised tissue). The scales however lacked the peg-and-socket articulations found in some other groups. The skulls of megalichthyids had a rather low and broad shape. Like the scales, the dermal bones of the skull bore a layer of cosmine. The nostrils were somewhat elongated, slit-like holes that were bordered in the front by the lateral rostral bone ...
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Ectosteorhachis
''Ectosteorhachis'' is an extinct genus of freshwater megalichthyid tetrapodomorphs that inhabited what is now North America during the Permian period (Cisuralian epoch, about 299 to 272 million years ago). It is the only known member of the subfamily Ectosteorhachinae''.'' Fossil remains are known from the United States. It contains a single species, ''E. nitidus'' from the Asselian/Sakmarian to Kungurian of Oklahoma and Utah (Hennesey and Cutler Formations). A second species, ''E. ciceronius'' Cope, 1883 from the Garber Formation of Oklahoma has no type specimen and is thus invalid. Remains tentatively assigned to ''Ectosteorhachis'' are known from the Washington Formation of Ohio Ohio ( ) is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. It borders Lake Erie to the north, Pennsylvania to the east, West Virginia to the southeast, Kentucky to the southwest, Indiana to the .... References Megalichthyidae Cisuralian ...
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Sarcopterygian
Sarcopterygii (; )—sometimes considered synonymous with Crossopterygii ()—is a clade (traditionally a class or subclass) of vertebrate animals which includes a group of bony fish commonly referred to as lobe-finned fish. These vertebrates are characterised by prominent muscular limb buds (lobes) within their fins, which are supported by articulated appendicular skeletons. This is in contrast to the other clade of bony fish, the Actinopterygii, which have only skin-covered bony spines supporting the fins. The tetrapods, a mostly terrestrial clade of vertebrates, are now recognized as having evolved from sarcopterygian ancestors and are most closely related to lungfishes. Their paired pectoral and pelvic fins evolved into limbs, and their foregut diverticulum eventually evolved into air-breathing lungs. Cladistically, this would make the tetrapods a subgroup within Sarcopterygii and thus sarcopterygians themselves. As a result, the phrase "lobe-finned fish" normally re ...
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Cladarosymblema
''Cladarosymblema'' is an extinct genus of megalichthyid tetrapodomorphs which lived in Australia during the Early Carboniferous Early may refer to: Places in the United States * Early, Iowa, a city * Early, Texas, a city * Early Branch, a stream in Missouri * Early County, Georgia * Fort Early, Georgia, an early 19th century fort Music * Early B, stage name of Jamaican d .... References Clement AM, Cloutier R, Lu J, Perilli E, Maksimenko A, Long J. 2021. A fresh look at Cladarosymblema narrienense, a tetrapodomorph fish (Sarcopterygii: Megalichthyidae) from the Carboniferous of Australia, illuminated via X-ray tomography. PeerJ 9:e12597 https://doi.org/10.7717/peerj.12597 Megalichthyidae Prehistoric lobe-finned fish genera Mississippian sarcopterygians Carboniferous sarcopterygians of Australia {{paleo-tetrapodomorph-stub ...
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Mahalalepis
''Mahalalepis'' is an extinct genus of megalichthyid tetrapodomorphs which lived during the Devonian period The Devonian ( ) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago ( Ma), to the beginning of the succeeding C .... References Megalichthyidae Middle Devonian sarcopterygians Late Devonian sarcopterygians Devonian sarcopterygians of Antarctica Fossil taxa described in 1992 {{paleo-tetrapodomorph-stub ...
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Dermal Bone
A dermal bone or investing bone or membrane bone is a bony structure derived from intramembranous ossification forming components of the vertebrate skeleton, including much of the skull, jaws, gill covers, shoulder girdle, fin rays ( lepidotrichia), and the shells of turtles and armadillos. In contrast to endochondral bone, dermal bone does not form from cartilage that then calcifies, and it is often ornamented. Dermal bone is formed within the dermis and grows by accretion only – the outer portion of the bone is deposited by osteoblasts. The function of some dermal bone is conserved throughout vertebrates, although there is variation in shape and in the number of bones in the skull roof and postcranial structures. In bony fish, dermal bone is found in the fin rays and scales. A special example of dermal bone is the clavicle The clavicle, collarbone, or keybone is a slender, S-shaped long bone approximately long that serves as a strut between the scapula, shoulder b ...
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Freshwater
Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include non-salty mineral-rich waters, such as chalybeate springs. Fresh water may encompass frozen and meltwater in ice sheets, ice caps, glaciers, snowfields and icebergs, natural precipitations such as rainfall, snowfall, hail/ sleet and graupel, and surface runoffs that form inland bodies of water such as wetlands, ponds, lakes, rivers, streams, as well as groundwater contained in aquifers, subterranean rivers and lakes. Water is critical to the survival of all living organisms. Many organisms can thrive on salt water, but the great majority of vascular plants and most insects, amphibians, reptiles, mammals and birds need fresh water to survive. Fresh water is the water resource that is of the most and immediate use to humans. Fresh water is n ...
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Fish Scale
A fish scale is a small rigid plate that grows out of the skin of a fish. The skin of most jawed fishes is covered with these protective scales, which can also provide effective camouflage through the use of reflection and colouration, as well as possible hydrodynamic advantages. The term ''scale'' derives from the Old French , meaning a shell pod or husk. Scales vary enormously in size, shape, structure, and extent, ranging from strong and rigid armour plates in fishes such as shrimpfishes and boxfishes, to microscopic or absent in fishes such as eels and anglerfishes. The morphology (biology), morphology of a scale can be used to identify the species of fish it came from. Scales originated within the jawless ostracoderms, ancestors to all jawed fishes today. Most bony fishes are covered with the cycloid scales of salmon and carp, or the ctenoid scales of perch, or the ganoid scales of sturgeons and gars. Cartilaginous fishes (sharks and rays) are covered with placoi ...
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Cosmine
Cosmine is a spongy, bony material that makes up the dentine-like layers in the scales of the lobe-finned fishes of the class Sarcopterygii. Fish scales that include layers of cosmine are known as cosmoid scales. Description As traditionally described, cosmine consists of a layer of dentine covered by a continuous sheet of enamel. Pulp cavities, which secrete dentine tubules, are surrounded by a complex polygonal network of 'pore cavities' which pierce the overlying enamel layer, giving cosmine its characteristic dotted appearance. The pulp cavities and pore chambers are connected by a complex, reticulated pore canal network which continues into a layer of vascular bone beneath the dentine. The exact configuration of the pore canal network and shape of the pore chambers differs between various taxa, although the general organization into a single layer of enamel over dentine with pore canals with vascular bone underneath remains consistent, at least within the Sarcopterygii. ...
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Middle Devonian
In the geological timescale, the Middle Devonian epoch (from 393.3 ± 1.2 million years ago to 382.7 ± 1.6 million years ago) occurred during the Devonian period, after the end of the Emsian age. The Middle Devonian epoch is subdivided into two stages: Eifelian and Givetian. Life in the Middle Devonian In the middle Devonian the armored jawless fish known as ostracoderms were declining in diversity and instead the jawed fish were thriving and increasing in diversity in both the oceans and freshwater. The shallow, warm, oxygen-depleted waters of Devonian inland lakes, surrounded by primitive plants, provided the environment necessary for certain early fish to develop essential characteristics such as well developed lungs, ability to crawl out of the water and onto the land for short periods of time, possibly in search of food which would be developed by the tetrapods later in the Late Devonian which are descendents of these early fish. The earliest forest grew in the Middle De ...
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Nostril
A nostril (or naris , : nares ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbinates, whose function is to warm air on inhalation and remove moisture on exhalation. Fish do not breathe through noses, but they do have two small holes used for smelling, which can also be referred to as nostrils (with the exception of Cyclostomi, which have just one nostril). In humans, the nasal cycle is the normal ultradian cycle of each nostril's blood vessels becoming engorged in swelling, then shrinking. The nostrils are separated by the septum. The septum can sometimes be deviated, causing one nostril to appear larger than the other. With extreme damage to the septum and columella, the two nostrils are no longer separated and form a single larger external opening. Like other tetrapod A tetrapod (; from Ancient Greek :wikti ...
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Early Permian
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