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Lophotrochozoan
Lophotrochozoa (, "crest/wheel animals") is a clade of protostome animals within the Spiralia. The taxon was established as a monophyletic group based on molecular evidence. The clade includes animals like annelids, molluscs, bryozoans, brachiopods, and platyhelminthes. Groups Lophotrochozoa was defined in 1995 as the "last common ancestor of the three traditional lophophorate taxa (brachiopods, bryozoans, and phoronid worms), the mollusks and the annelids, and all of the descendants of that common ancestor". It is a cladistic definition (a node-based name), so the affiliation to Lophotrochozoa of spiralian groups not mentioned directly in the definition depends on the topology of the spiralian tree of life, and in some phylogenetic hypotheses, Lophotrochozoa may even be synonymous to Spiralia. Nemertea and Orthonectida (if not directly considered as part of Annelida) are probably lophotrochozoan phyla; Dicyemida, Gastrotricha, and Platyhelminthes may be lophotrochozoans or ...
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Brachiopod
Brachiopods (), phylum Brachiopoda, are a phylum of trochozoan animals that have hard "valves" (shells) on the upper and lower surfaces, unlike the left and right arrangement in bivalve molluscs. Brachiopod valves are hinged at the rear end, while the front can be opened for feeding or closed for protection. Two major categories are traditionally recognized, articulate and inarticulate brachiopods. The word "articulate" is used to describe the tooth-and-groove structures of the valve-hinge which is present in the articulate group, and absent from the inarticulate group. This is the leading diagnostic skeletal feature, by which the two main groups can be readily distinguished as fossils. Articulate brachiopods have toothed hinges and simple, vertically-oriented opening and closing muscles. Conversely, inarticulate brachiopods have weak, untoothed hinges and a more complex system of vertical and oblique (diagonal) muscles used to keep the two valves aligned. In many brachiopods, a ...
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Oikozetetes
The halkieriids are a group of fossil organisms from the Lower to Middle Cambrian. Their eponymous genus is ''Halkieria'' , which has been found on almost every continent in Lower to Mid Cambrian deposits, forming a large component of the small shelly fossil assemblages. The best known species is ''Halkieria evangelista'', from the North Greenland Sirius Passet Lagerstätte, in which complete specimens were collected on an expedition in 1989. The fossils were described by Simon Conway Morris and John Peel in a short paper in 1990 in the journal ''Nature''. Later a more thorough description was undertaken in 1995 in the journal ''Philosophical Transactions of the Royal Society of London'' and wider evolutionary implications were posed. The group is sometimes equated to Sachitida, although as originally envisaged, this group includes the wiwaxiids and is thus equivalent to the Halwaxiida. History of discovery Armor plates called "sclerites" had long been known as elements of th ...
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Spiralia
The Spiralia are a morphologically diverse clade of protostome animals, including within their number the molluscs, annelids, platyhelminths and other taxa. The term ''Spiralia'' is applied to those phyla that exhibit canonical spiral cleavage, a pattern of early development found in most (but not all) members of the Lophotrochozoa. Distribution of spiralian development across phylogeny Members of the molluscs, annelids, platyhelminths and nemerteans have all been shown to exhibit spiral cleavage in its classical form. Other spiralian phyla ( rotifers, brachiopods, phoronids, gastrotrichs, and bryozoans) are also said to display a derived form of spiral cleavage in at least a portion of their constituent species, although evidence for this is sparse. Lophotrochozoa within Spiralia Previously, spiral cleavage was thought to be unique to the Spiralia in the strictest sense—animals such as molluscs and annelids which exhibit classical spiral cleavage. The presence of s ...
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Spiralia
The Spiralia are a morphologically diverse clade of protostome animals, including within their number the molluscs, annelids, platyhelminths and other taxa. The term ''Spiralia'' is applied to those phyla that exhibit canonical spiral cleavage, a pattern of early development found in most (but not all) members of the Lophotrochozoa. Distribution of spiralian development across phylogeny Members of the molluscs, annelids, platyhelminths and nemerteans have all been shown to exhibit spiral cleavage in its classical form. Other spiralian phyla ( rotifers, brachiopods, phoronids, gastrotrichs, and bryozoans) are also said to display a derived form of spiral cleavage in at least a portion of their constituent species, although evidence for this is sparse. Lophotrochozoa within Spiralia Previously, spiral cleavage was thought to be unique to the Spiralia in the strictest sense—animals such as molluscs and annelids which exhibit classical spiral cleavage. The presence of s ...
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Nemertea
Nemertea is a phylum of animals also known as ribbon worms or proboscis worms, consisting of 1300 known species. Most ribbon worms are very slim, usually only a few millimeters wide, although a few have relatively short but wide bodies. Many have patterns of yellow, orange, red and green coloration. The foregut, stomach and intestine run a little below the midline of the body, the anus is at the tip of the tail, and the mouth is under the front. A little above the gut is the rhynchocoel, a cavity which mostly runs above the midline and ends a little short of the rear of the body. All species have a proboscis which lies in the rhynchocoel when inactive but everts to emerge just above the mouth to capture the animal's prey with venom. A highly extensible muscle in the back of the rhynchocoel pulls the proboscis in when an attack ends. A few species with stubby bodies filter feed and have suckers at the front and back ends, with which they attach to a host. The brain is a rin ...
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Bryozoa
Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869living species are known. At least two genera are solitary (''Aethozooides'' and ''Monobryozoon''); the rest are colonial. The terms Polyzoa and Bryozoa were introduced in 1830 and 1831, respectively. Soon after it was named, another group of animals was discovered whose filtering mechanism looked similar, so it was included in Bryozoa until 1869, when the two groups wer ...
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Annelid
The annelids (Annelida , from Latin ', "little ring"), also known as the segmented worms, are a large phylum, with over 22,000 extant species including ragworms, earthworms, and leeches. The species exist in and have adapted to various ecologies – some in marine environments as distinct as tidal zones and hydrothermal vents, others in fresh water, and yet others in moist terrestrial environments. The Annelids are bilaterally symmetrical, triploblastic, coelomate, invertebrate organisms. They also have parapodia for locomotion. Most textbooks still use the traditional division into polychaetes (almost all marine), oligochaetes (which include earthworms) and leech-like species. Cladistic research since 1997 has radically changed this scheme, viewing leeches as a sub-group of oligochaetes and oligochaetes as a sub-group of polychaetes. In addition, the Pogonophora, Echiura and Sipuncula, previously regarded as separate phyla, are now regarded as sub-groups of polychae ...
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Hyolitha
Hyoliths are animals with small conical shells, known as fossils from the Palaeozoic era. They are at least considered as lophotrochozoan, and possibly being lophophorates, a group which includes the brachiopods, while others consider them as being basal lophotrochozoans, or even molluscs. Morphology The shell of a hyolith is typically one to four centimeters in length, triangular or elliptical in cross section. Some species have rings or stripes. It comprises two parts: the main conical shell (previously referred to as a ‘conch’) and a cap-like operculum. Some also had two curved supports known as ''helens'' They are calcareous – probably aragonitic All of these structures grew by marginal accretion. Shell microstructure The orthothecid shell has an internal layer with a microstructure of transverse bundles, and an external layer comprising longitudinal bundles. Helens Some hyoliths had helens, long structures that taper as they coil gently in a logarithmic spira ...
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Macrodontophion
''Macrodontophion'' (meaning "long-toothed snake") is the name given to a dubious genus of Lophotrochozoa, lophotrochozoan from the Early Devonian Dniester Series of Podolia, Ukraine. It was described by Adalbert Zborzewsky in 1834, but was never given a species epithet, and is considered a ''nomen dubium'', because it is based only on fragments, such as the holotype, a shell of . The known specimens of ''Macrodontophion'' are presumed lost. Several of the specimens Zborzewski described are listed as being in his private collection, while others were said to be held by his colleagues. Taxonomy ''Macrodontophion'' was originally believed to have been a snake tooth that belonged to an animal similar to ''Ophisaurus'', or the cephalopod ''Beloptera'', by Zborzewsky in 1834. Since ''Megalosaurus'' was mentioned in the same paragraph that the genus name ''Macrodontophion'' first appears, many palaeontologists, such as Alfred Romer, Romer in 1956, Steel in 1970 and Romer again in 1976, ...
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Mollusca
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is estimated between 60,000 and 100,000 additional species. The proportion of undescribed species is very high. Many taxa remain poorly studied. Molluscs are the largest marine phylum, comprising about 23% of all the named marine organisms. Numerous molluscs also live in freshwater and terrestrial habitats. They are highly diverse, not just in size and anatomical structure, but also in behaviour and habitat. The phylum is typically divided into 7 or 8  taxonomic classes, of which two are entirely extinct. Cephalopod molluscs, such as squid, cuttlefish, and octopuses, are among the most neurologically advanced of all invertebrates—and either the giant squid or the colossal squid is the largest known invertebrate species. The ...
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Lophophorata
The Lophophorata are a Lophotrochozoan clade consisting of the Brachiozoa and the Bryozoa. They have a lophophore. Molecular phylogenetic analyses suggest that lophophorates are protostomes, but on morphological grounds they have been assessed as deuterostome Deuterostomia (; in Greek) are animals typically characterized by their anus forming before their mouth during embryonic development. The group's sister clade is Protostomia, animals whose digestive tract development is more varied. Some ...s. Fossil finds of a segmented worm named Wufengella suggest that they evolved from a worm close to annelids. References {{Protostome-stub ...
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Bryozoa
Bryozoa (also known as the Polyzoa, Ectoprocta or commonly as moss animals) are a phylum of simple, aquatic invertebrate animals, nearly all living in sedentary colonies. Typically about long, they have a special feeding structure called a lophophore, a "crown" of tentacles used for filter feeding. Most marine bryozoans live in tropical waters, but a few are found in oceanic trenches and polar waters. The bryozoans are classified as the marine bryozoans (Stenolaemata), freshwater bryozoans (Phylactolaemata), and mostly-marine bryozoans (Gymnolaemata), a few members of which prefer brackish water. 5,869living species are known. At least two genera are solitary (''Aethozooides'' and ''Monobryozoon''); the rest are colonial. The terms Polyzoa and Bryozoa were introduced in 1830 and 1831, respectively. Soon after it was named, another group of animals was discovered whose filtering mechanism looked similar, so it was included in Bryozoa until 1869, when the two groups wer ...
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