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Fagales
The Fagales are an order of flowering plants in the rosid group of dicotyledons, including some of the best-known trees. Well-known members of Fagales include: beeches, chestnuts, oaks, walnut, pecan, hickory, birches, alders, hazels, hornbeams, she-oaks, and southern beeches. The order name is derived from genus ''Fagus'' (beeches). Systematics Fagales include the following seven families, according to the APG III system of classification: * Betulaceae – birch family ('' Alnus'', '' Betula'', ''Carpinus'', '' Corylus'', ''Ostrya'', and '' Ostryopsis'') *Casuarinaceae – she-oak family ('' Allocasuarina'', '' Casuarina'', ''Ceuthostoma'', and '' Gymnostoma'') *Fagaceae – beech family ('' Castanea'', '' Castanopsis'', '' Chrysolepis'', ''Fagus'', '' Lithocarpus'', '' Notholithocarpus'', ''Quercus'', and '' Trigonobalanus'') *Juglandaceae – walnut family (''Alfaroa'', '' Carya'', '' Cyclocarya'', '' Engelhardia'', ''Juglans'', '' Oreomunnea'', '' Platycarya'', '' P ...
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APG III System
The APG III system of flowering plant classification is the third version of a modern, mostly molecular-based, system of plant taxonomy being developed by the Angiosperm Phylogeny Group (APG). Published in 2009, it was superseded in 2016 by a further revision, the APG IV system. Along with the publication outlining the new system, there were two accompanying publications in the same issue of the Botanical Journal of the Linnean Society: * The first, by Chase & Reveal, was a formal phylogenetic classification of all land plants (embryophytes), compatible with the APG III classification. As the APG have chosen to eschew ranks above order, this paper was meant to fit the system into the existing Linnaean hierarchy for those that prefer such a classification. The result was that all land plants were placed in the class Equisetopsida, which was then divided into 16 subclasses and a multitude of superorders. * The second, by Haston ''et al.'', was a linear sequence of families fol ...
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Ticodendraceae
''Ticodendron incognitum'' is the only species of ''Ticodendron'', and the only member of the family Ticodendraceae. It is most closely related to the family Betulaceae. It was discovered only in 1989 in Costa Rica, having been overlooked previously due to its habitat in poorly researched cloud forests and its very 'ordinary' appearance; further research showed its range extends from southern Mexico (Veracruz, Oaxaca, Chiapas), south through Central America to Panama. It is a tree, 20–30 m tall, superficially resembling an alder in appearance, with alternate, simple leaves 5–12 cm long with a serrated margin. It is generally dioecious, with separate male and female plants. Fossil record †''Ferrignocarpus bivalvis'' fossil fruits, from the Middle Eocene of Oregon and the Early Eocene London Clay flora of southern England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of w ...
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Allocasuarina
''Allocasuarina'', commonly known as sheoak or she-oak, is a genus of flowering plants in the family Casuarinaceae and is endemic to Australia. Plants in the genus ''Allocasuarina'' are trees or shrubs with soft, pendulous, green branchlets, the leaves reduced to scale-like teeth. Allocasuarinas are either monoecious or dioecious, the flowers never bisexual. Male and female flowers are arranged in spikes, the female spikes developing into cone-like structures enclosing winged seeds. The genera ''Allocasuarina'' and '' Casuarina'' are similar, and many formerly in the latter now included in ''Allocasuarina''. Description Plants in the genus ''Allocasuarina'' are trees or shrubs with soft, pendulous, green branchlets, the leaves reduced to 4 to 14 scale-like teeth arranged around in whorls around ribbed, jointed branchlets. Allocasuarinas have separate male and female flowers, sometimes on one plant (monoecious), otherwise on separate male and female plants, (dioecious). Male fl ...
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Dicotyledon
The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, that the seed has two embryonic leaves or cotyledons. There are around 200,000 species within this group. The other group of flowering plants were called monocotyledons (or monocots), typically each having one cotyledon. Historically, these two groups formed the two divisions of the flowering plants. Largely from the 1990s onwards, molecular phylogenetic research confirmed what had already been suspected: that dicotyledons are not a group made up of all the descendants of a common ancestor (i.e., they are not a monophyletic group). Rather, a number of lineages, such as the magnoliids and groups now collectively known as the basal angiosperms, diverged earlier than the monocots did; in other words, monocots evolved from within the ...
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Rosid
The rosids are members of a large clade (monophyletic group) of flowering plants, containing about 70,000 species, more than a quarter of all angiosperms. The clade is divided into 16 to 20 orders, depending upon circumscription and classification. These orders, in turn, together comprise about 140 families. Fossil rosids are known from the Cretaceous period. Molecular clock estimates indicate that the rosids may have originated in the Aptian or Albian stages of the Cretaceous, between 125 and 99.6 million years ago. Today's broadleaved forests are dominated by rosid species, which in turn help with diversification in many other living lineages. Additionally, rosid herbs and shrubs are a significant part of arctic/alpine and temperate floras. The clade also includes some aquatic, desert and parasitic plants. Name The name is based upon the name " Rosidae", which had usually been understood to be a subclass. In 1967, Armen Takhtajan showed that the correct basis for the name "R ...
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Hickory
Hickory is a common name for trees composing the genus ''Carya'', which includes 19 species accepted by ''Plants of the World Online''. Seven species are native to southeast Asia in China, Indochina, and northeastern India (Assam), and twelve are native to North America. A number of hickory species are used for their edible nuts or for their wood. Etymology The name "hickory" derives from a Native American languages, Native American word in an Algonquian languages, Algonquian language (perhaps Powhatan language, Powhatan). It is a shortening of ''pockerchicory'', ''pocohicora'', or a similar word, which may be the name for the hickory tree's nut, or may be a plant milk, milky drink made from such nuts. The genus name ''Carya'' is , ''káryon'', meaning "nut (fruit), nut". Description Hickories are temperate forest, temperate to tropical and subtropical moist broadleaf forests, subtropical forest trees with pinnation, pinnately compound leaves and large nut (fruit), nuts. Most ...
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Myricaceae
Myricaceae is a small family of dicotyledonous shrubs and small trees in the order Fagales with its type genus ''Myrica'', the sweet gales. There are three genera in the family, although some botanists separate many species from Myrica into a fourth genus ''Morella''. About 55 species are usually accepted in ''Myrica'' (with ''Morella'' included), one in '' Canacomyrica'', and one in '' Comptonia''. Well-known members of this family include bayberry and sweetfern. * '' Canacomyrica'' Guillaumin 1940 * '' Comptonia'' L'Hér. ex Aiton 1789 * ''Myrica'' L. 1753 (includes: '' Morella'' Lour. 1790) Systematics Modern molecular phylogenetics Molecular phylogenetics () is the branch of phylogeny that analyzes genetic, hereditary molecular differences, predominantly in DNA sequences, to gain information on an organism's evolutionary relationships. From these analyses, it is possible to ... suggest the following relationships: References Rosid families {{Faga ...
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Nothofagaceae
''Nothofagus'', also known as the southern beeches, is a genus of 43 species of trees and shrubs native to the Southern Hemisphere, found across southern South America (Chile, Argentina) and east and southeast Australia, New Zealand, New Guinea, and New Caledonia. The species are ecological dominants in many temperate forests in these regions. Some species are reportedly naturalised in Germany and Great Britain. The genus has a rich fossil record of leaves, cupules, and pollen, with fossils extending into the late Cretaceous period and occurring in Australia, New Zealand, Antarctica, and South America. Description The leaves are toothed or entire, evergreen or deciduous. The fruit is a small, flattened or triangular nut, borne in cupules containing one to seven nuts. Reproduction Many individual trees are extremely old, and at one time, some populations were thought to be unable to reproduce in present-day conditions where they were growing, except by suckering ( clonal re ...
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Casuarinaceae
The Casuarinaceae are a family of dicotyledonous flowering plants placed in the order Fagales, consisting of four genera and 91 species of trees and shrubs native to eastern Africa, Australia, Southeast Asia, Malesia, Papuasia, and the Pacific Islands. At one time, all species were placed in the genus '' Casuarina''. Lawrence Alexander Sidney Johnson separated out many of those species and renamed them into the new genera of '' Gymnostoma'' in 1980 and 1982, '' Allocasuarina'' in 1982, and ''Ceuthostoma'' in 1988, with some additional formal descriptions of new species in each other genus. At the time, it was somewhat controversial. The monophyly of these genera was later supported in a 2003 phylogenetic study of the family. In the Wettstein system, this family was the only one placed in the order Verticillatae. Likewise, in the Engler, Cronquist, and Kubitzki systems, the Casuarinaceae were the only family placed in the order Casuarinales. Members of this family are charac ...
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Ostryopsis
''Ostryopsis'' is a small genus of deciduous shrubs belonging to the birch family Betulaceae. The species have no common English name, though hazel-hornbeam has been suggested, reflecting their similarities to the closely related hazels and hop-hornbeams. The genus is native to China. They are shrubs reaching 3–5 m tall, with alternate, double-toothed hazel-like leaves 2–7 cm long. The flowers are produced in spring, with separate male and female catkins. The fruit form in clusters 3–5 cm long with 6-10 seeds; each seed is a small nut 4–6 mm long, fully enclosed in a sheath-like involucre. The local people in Northeast China has found hazelnut The hazelnut is the fruit of the hazel tree and therefore includes any of the nuts deriving from species of the genus '' Corylus'', especially the nuts of the species ''Corylus avellana''. They are also known as cobnuts or filberts according to ...s of '' Ostryopsis davidiana'' and '' Corylus mandshurica'' are ...
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Flowering Plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (). The term angiosperm is derived from the Ancient Greek, Greek words (; 'container, vessel') and (; 'seed'), meaning that the seeds are enclosed within a fruit. The group was formerly called Magnoliophyta. Angiosperms are by far the most diverse group of Embryophyte, land plants with 64 Order (biology), orders, 416 Family (biology), families, approximately 13,000 known Genus, genera and 300,000 known species. They include all forbs (flowering plants without a woody Plant stem, stem), grasses and grass-like plants, a vast majority of broad-leaved trees, shrubs and vines, and most aquatic plants. Angiosperms are distinguished from the other major seed plant clade, the gymnosperms, by having flowers, xylem consisting of vessel elements instead of tracheids, endosperm within their seeds, and fruits that completely envelop the seeds. The ancestors of flowering plants diverged from the commo ...
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Juglandaceae
The Juglandaceae are a plant family known as the walnut family. They are trees, or sometimes shrubs, in the order Fagales. Members of this family are native to the Americas, Eurasia, and Southeast Asia. The nine or ten genera in the family have a total of around 50 species, and include the commercially important nut-producing trees walnut (''Juglans''), pecan (''Carya illinoinensis''), and hickory (''Carya''). The Persian walnut, '' Juglans regia'', is one of the major nut crops of the world. Walnut, hickory, and gaulin are also valuable timber trees while pecan wood is also valued as cooking fuel. Description Members of the walnut family have large, aromatic leaves that are usually alternate, but opposite in '' Alfaroa'' and '' Oreomunnea''. The leaves are pinnately compound or ternate, and usually 20–100 cm long. The trees are wind-pollinated, and the flowers are usually arranged in catkins. The fruits of the Juglandaceae are often confused with drupes but are ac ...
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