Eutheriodonts
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Eutheriodonts
Eutheriodontia is a clade of therapsids which appear during the Middle Permian and which includes therocephalians and cynodonts, this latter group including mammals and related forms. With the dicynodonts, they form one of two lineages of therapsids that survived the End-Permian extinction and which diversified again during the Triassic, before the majority of them disappeared before or during the Triassic–Jurassic extinction, except for a lineage of cynodonts that later gave rise to mammals. Classification The clade was named in 1986 by James Allen Hopson and Herbert Richard Barghusen, the name meaning the "true Theriodontia". Within Hopson's system, the Eutheriodontia are the sister group of the Gorgonopsia within the Theriodontia. A close relationship between therocephalians and cynodonts had been recognized for many years. In 2001 the Eutheriodontia were defined as the least inclusive clade including Mammalia and ''Bauria''.James A. Hopson and James W. Kitching, 20 ...
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Gorgonopsia
Gorgonopsia (from the Greek Gorgon, a mythological beast, and 'aspect') is an extinct clade of Saber-toothed predator, sabre-toothed therapsids from the Middle Permian, Middle to the Upper Permian, roughly between 270 and 252 million years ago. They are characterised by a long and narrow skull, as well as elongated upper and sometimes lower canine teeth and incisors which were likely used as slashing and stabbing weapons. Postcanine teeth are generally reduced or absent. For hunting large prey, they possibly used a bite-and-retreat tactic, ambushing and taking a debilitating bite out of the target, and following it at a safe distance before its injuries exhausted it, whereupon the gorgonopsian would grapple the animal and deliver a killing bite. They would have had an exorbitant gape, possibly in excess of 90°, without having to unhinge the jaw. They markedly increased in size as time went on, growing from small skull lengths of in the Middle Permian to bear-like proportions of ...
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Theriodontia
The theriodonts (clade Theriodontia) are a major group of therapsids which appeared during the Middle Permian and which includes the gorgonopsians and the eutheriodonts, itself including the therocephalians and the cynodonts. Naming In 1876, Richard Owen named a suborder Theriodontia, which he divided into the Cynodontia and the Gomphodontia. In 1956, D. M. S. Watson and Alfred Romer divided Therapsida into Theriodontia, which predominantly included carnivorous species, and the mostly herbivorous Anomodontia, defined as comprising Dinocephalia as well as Venyukovioidea and Dicynodontia. The modern clade concept was devised by James Allen Hopson. In his system, Theriodontia fall into two main groups: the Gorgonopsia and the Eutheriodontia. The latter consist of the Therocephalia and Cynodontia.J.A. Hopson. 1999. "Therapsids". ''Encyclopedia of Paleontology'' 2: 1256-1266 Evolution Theriodonts appeared at the same time as their sister group within the Neotherapsida, the ...
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Permian–Triassic Extinction Event
The Permian–Triassic extinction event (also known as the P–T extinction event, the Late Permian extinction event, the Latest Permian extinction event, the End-Permian extinction event, and colloquially as the Great Dying,) was an extinction event that occurred approximately 251.9 Million years ago, million years ago (mya), at the boundary between the Permian and Triassic geologic periods, and with them the Paleozoic and Mesozoic eras. It is Earth's most severe known extinction event, with the extinction of 57% of Family (biology), biological families, 62% of genera and 81% of marine biology, marine species, (previous estimates of 90–96% marine species extinction were due to historical conflation with the End-Capitanian mass extinction event, end-Capitanian mass extinction which occurred 7–10 million years earlier) and 70% of terrestrial vertebrate species. It is also the greatest known mass extinction of insects. It is the greatest of the "Big Five" mass extinctions of t ...
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Middle Permian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian is also known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. ...
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Prozostrodontia
Prozostrodontia is a clade of cynodonts including mammaliaforms and their closest relatives such as Tritheledontidae and Tritylodontidae. It was erected as a node-based taxon by Liu and Olsen (2010) and defined as the least inclusive clade containing ''Prozostrodon brasiliensis'', ''Tritylodon langaevus'', ''Pachygenelus monus'', and ''Mus musculus'' (the house mouse). Prozostrodontia is diagnosed by several characters, including: * Reduced prefrontal bone, prefrontal and postorbital bones, with the disappearance of a strut of bone called the postorbital bar separating the Orbit (anatomy), eye socket from the Temporal fenestra, temporal region * Unfused Mandibular symphysis, symphysis between the dentary bones in the lower jaw * The presence of a small hole within the eye socket called the sphenopalatine foramen * A long sagittal crest extending to the rearmost part of the lambdoid suture, lambdoidal crest at the back of the skull * Neural spines of the dorsal vertebrae angled back ...
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Skull Roof
The skull roof or the roofing bones of the skull are a set of bones covering the brain, eyes and nostrils in bony fishes, including land-living vertebrates. The bones are derived from dermal bone and are part of the dermatocranium. In comparative anatomy, the term is applied to the whole dermatocranium. Romer, A.S. & T.S. Parsons. 1977. ''The Vertebrate Body.'' 5th ed. Saunders, Philadelphia. (6th ed. 1985) In general anatomy, the roofing bones may refer specifically to the bones that form above and alongside the brain and neurocranium (i.e., excluding the marginal upper jaw bones such as the maxilla and premaxilla). In human anatomy, the skull roof often refers specifically to the skullcap. Origin Early armoured fish (such as jawless ostracoderms and jawed placoderms) did not have a skull in the common understanding of the word, but instead had a cartilaginous endocranium that was partially open from above. The loose cartilage was topped by dermal bones forming armo ...
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Alisphenoid
The greater wing of the sphenoid bone, or alisphenoid, is a bony process of the sphenoid bone, positioned in the skull behind each eye. There is one on each side, extending from the side of the body of the sphenoid and curving upward, laterally, and backward. Structure The greater wings of the sphenoid are two strong processes of bone, which arise from the sides of the body, and are curved upward, laterally, and backward; the posterior part of each projects as a triangular process that fits into the angle between the squamous and the petrous part of the temporal bone and presents at its apex a downward-directed process, the spine of sphenoid bone. Cerebral surface The superior or cerebral surface of each greater wing ig. 1forms part of the middle cranial fossa; it is deeply concave, and presents depressions for the convolutions of the temporal lobe of the brain. It has a number of foramina (holes) in it: * The foramen rotundum is a circular aperture at its anterior and medial ...
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Palate
The palate () is the roof of the mouth in humans and other mammals. It separates the oral cavity from the nasal cavity. A similar structure is found in crocodilians, but in most other tetrapods, the oral and nasal cavities are not truly separated. The palate is divided into two parts, the anterior, bony hard palate and the posterior, fleshy soft palate (or velum). Structure Innervation The maxillary nerve branch of the trigeminal nerve supplies sensory innervation to the palate. Development The hard palate forms before birth. Variation If the fusion is incomplete, a cleft palate results. Function in humans When functioning in conjunction with other parts of the mouth, the palate produces certain sounds, particularly velar, palatal, palatalized, postalveolar, alveolopalatal, and uvular consonants. History Etymology The English synonyms palate and palatum, and also the related adjective palatine (as in palatine bone), are all from the Latin ''palatum' ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel Phenotypic trait, character or character state that has evolution, evolved from its ancestral form (or Plesiomorphy and symplesiomorphy, plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxon, taxa and is therefore Hypothesis#Scientific hypothesis, hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies Homology (biology), homology. Examples of apomorphy are the presence of Terrestrial locomotion#Posture, erect gait, fur, Evolution of mammalian auditory ossicles, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a Terrestrial locomotion#Posture, sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word ...
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Postorbital Bar
The postorbital bar (or postorbital bone) is a bony arched structure that connects the frontal bone of the skull to the zygomatic arch, which runs laterally around the eye socket. It is a trait that only occurs in mammalian taxa, such as most strepsirrhine primates and the hyrax, while haplorhine primates have evolved fully enclosed sockets. One theory for this evolutionary difference is the relative importance of vision to both orders. As haplorrhines (tarsiers and simians) tend to be diurnal, and rely heavily on visual input, many strepsirrhines are nocturnal and have a decreased reliance on visual input. Postorbital bars evolved several times independently during mammalian evolution and the evolutionary histories of several other clades. Some species, such as tarsiers, have a postorbital septum. This septum can be considered as joined processes with a small articulation between the frontal bone, the zygomatic bone and the alisphenoid bone and is therefore different from the p ...
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Secondary Palate
The secondary palate is an anatomical structure that divides the nasal cavity from the oral cavity in many vertebrates. In human embryology, it refers to that portion of the hard palate that is formed by the growth of the two palatine shelves medially and their mutual fusion in the midline. It forms the majority of the adult palate and meets the primary palate at the incisive foramen. Clinical significance Secondary palate development begins in the sixth week of pregnancy and can lead to cleft palate when development goes awry. There are three major mechanisms known to cause this failure: #Growth retardation — Palatal shelves do not grow enough to meet each other. #Mechanical obstruction — Improper mouth size, or abnormal anatomical structures in the embryonic mouth prevent fully grown shelves from meeting each other. #Midline epithelial dysfunction (MED) - The surface mucosa of embryonic shelves is impaired, which causes a failure of palatal fusion. Evolution T ...
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Bauria
''Bauria'' is an extinct genus of the suborder Therocephalia that existed during the Early Triassic, Early and Middle Triassic period, around 246-251 million years ago. It belonged to the family Bauriidae. ''Bauria'' was probably a herbivore or omnivore. It lived in South Africa, specifically in the Burgersdorp Formation in South Africa. Taxonomy ''Bauria'' was named by Robert Broom in 1909 and found at Winnaarsbaken, South Africa. The first species Broom discovered, ''Bauria cynops'', was a reasonably complete skull, but according to the first description somewhat poorly preserved, and apparently equally poorly prepared. Five other specimens were later found at different points in time, with mostly skulls being found. There have only been two known species of ''Bauria'' that have been discovered so far, with the first species, ''Bauria cynops'', being known from 6 different skulls in varying conditions of poor to excellent. The second species, ''Bauria robusta'' is known f ...
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