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Exaptation
Exaptation and the related term co-option describe a shift in the function of a trait during evolution. For example, a trait can evolve because it served one particular function, but subsequently it may come to serve another. Exaptations are common in both anatomy and behaviour. Bird feathers are a classic example. Initially they may have evolved for temperature regulation, but later were adapted for flight. When feathers were first used to aid in flight, that was an exaptive use. They have since then been shaped by natural selection to improve flight, so in their current state they are best regarded as adaptations for flight. So it is with many structures that initially took on a function as an exaptation: once molded for a new function, they become further adapted for that function. Interest in exaptation relates to both the process and products of evolution: the process that creates complex traits and the products (functions, anatomical structures, biochemicals, etc.) that may ...
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Elisabeth Vrba
Elisabeth S. Vrba (born 17 May 1942) is a paleontologist at Yale University who developed the turnover-pulse hypothesis. Education Vrba earned her Ph.D. in Zoology and Palaeontology at the University of Cape Town, in 1974. Vrba studied zoology and mathematical statistics at the University of Cape Town to earn her undergraduate degree. She remained there for doctoral study in zoology and paleontology to earn her Ph.D. After receiving her doctorate, Vrba conducted her early research on African fossil records over the last several million years, tracking the sequence of fossils from analyzing the geological strata and analyzing the morphology of the fossils. She was the chief assistant to Charles Kimberlin Brain during his directorship of the Transvaal Museum. Career She has been a faculty member at the Department of Geology & Geophysics, Yale University, since the early 1980s. She is well known for developing the turnover-pulse hypothesis, as well as coining the word exaptat ...
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Evolution
Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation tends to exist within any given population as a result of genetic mutation and recombination. Evolution occurs when evolutionary processes such as natural selection (including sexual selection) and genetic drift act on this variation, resulting in certain characteristics becoming more common or more rare within a population. The evolutionary pressures that determine whether a characteristic is common or rare within a population constantly change, resulting in a change in heritable characteristics arising over successive generations. It is this process of evolution that has given rise to biodiversity at every level of biological organisation, including the levels of species, individual organisms, and molecules. The theory of evol ...
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Jürgen Brosius
Jürgen Brosius (born 1948) in Saarbrücken) is a German molecular geneticist and evolutionary biologist. He was professor and director of the Institute of Experimental Pathology at the University of Münster. Some of his scientific contributions involve the first genetic sequencing of a ribosomal RNA operon, the design of plasmids for studying gene expression, expression vectors for high-level production of recombinant proteins and RNA, RNA biology, RNomics as well as the significance of retroposition for plasticity and evolution of genomes, genes and gene modules including regulatory sequences or elements. Biography Early life and education Brosius studied chemistry and pharmacy at the Goethe University of Frankfurt and in 1974 graduated and completed the Staatsexamen (state examination) in Pharmacy. Subsequently, he pursued his Ph.D. work in biochemistry and molecular biology at the Max Planck Institute for Molecular Genetics in Berlin Dahlem in which Heinz-Günter Wittmann ...
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Stephen Jay Gould
Stephen Jay Gould (; September 10, 1941 – May 20, 2002) was an American paleontologist, evolutionary biologist, and historian of science. He was one of the most influential and widely read authors of popular science of his generation. Gould spent most of his career teaching at Harvard University and working at the American Museum of Natural History in New York. In 1996, Gould was hired as the Vincent Astor Visiting Research Professor of Biology at New York University, after which he divided his time teaching between there and Harvard. Gould's most significant contribution to evolutionary biology was the theory of punctuated equilibrium developed with Niles Eldredge in 1972.Eldredge, Niles, and S. J. Gould (1972)"Punctuated equilibria: an alternative to phyletic gradualism."In T.J.M. Schopf, ed., ''Models in Paleobiology''. San Francisco: Freeman, Cooper and Company, pp. 82–115. The theory proposes that most evolution is characterized by long periods of evolutionary stabili ...
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Teleology In Biology
Teleology in biology is the use of the language of goal-directedness in accounts of evolutionary adaptation, which some biologists and philosophers of science find problematic. The term teleonomy has also been proposed. Before Darwin, organisms were seen as existing because God had designed and created them; their features such as eyes were taken by natural theology to have been made to enable them to carry out their functions, such as seeing. Evolutionary biologists often use similar teleological formulations that invoke purpose, but these imply natural selection rather than actual goals, whether conscious or not. Dissenting biologists and religious thinkers held that evolution itself was somehow goal-directed ( orthogenesis), and in vitalist versions, driven by a purposeful life force. Since such views are now discredited, with evolution working by natural selection acting on inherited variation, the use of teleology in biology has attracted criticism, and attempts have ...
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Zachary Blount
Zachary D. Blount is an American evolutionary biologist best known for his work on the evolution of a key innovation, aerobic growth on citrate, in one of the twelve populations of the ''E. coli'' long-term evolution experiment. Blount is a research assistant professor working with Richard Lenski at Michigan State University. He was previously a postdoctoral research assistant for Lenski, and was a visiting assistant professor of biology at Kenyon College from 2018 to 2019. Early life and education Zachary D. Blount was born and raised in Georgia where he developed an early interest in science. He earned high honors in Biology at Georgia Institute of Technology, where he was introduced to microbiology by professor emeritus Thomas Tornabene. He obtained his master's from the University of Cincinnati, where he worked on insertion sequence elements of hyperthermoacidophilic Archaea of the genus ''Sulfolobus'' under the guidance of Dennis Grogan. He moved to Michigan State Unive ...
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Evolution Of Mammalian Auditory Ossicles
The evolution of mammalian auditory ossicles was an evolutionary event that resulted in the formation of the bones of the mammalian middle ear. These bones, or ossicles, are a defining characteristic of all mammals. The event is well-documented and important as a demonstration of transitional forms and exaptation, the re-purposing of existing structures during evolution. The ossicles evolved from skull bones present in most tetrapods, including the reptilian lineage. The reptilian quadrate bone, articular bone, and columella evolved into the mammalian incus, malleus, and stapes (anvil, hammer, and stirrup), respectively. In reptiles, the eardrum is connected to the inner ear via a single bone, the columella, while the upper and lower jaws contain several bones not found in mammals. Over the course of the evolution of mammals, one bone from the lower and one from the upper jaw (the articular and quadrate bones) lost their purpose in the jaw joint and migrated to the middle ear ...
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Great Oxygenation Event
The Great Oxidation Event (GOE), also called the Great Oxygenation Event, the Oxygen Catastrophe, the Oxygen Revolution, the Oxygen Crisis, or the Oxygen Holocaust, was a time interval during the Paleoproterozoic era when the Earth's atmosphere and the shallow ocean first experienced a rise in the amount of oxygen. This began approximately 2.460–2.426 Ga (billion years) ago, during the Siderian period, and ended approximately 2.060 Ga, during the Rhyacian. Geological, isotopic, and chemical evidence suggests that biologically-produced molecular oxygen ( dioxygen, O2) started to accumulate in Earth's atmosphere and changed it from a weakly reducing atmosphere practically free of oxygen into an oxidizing atmosphere containing abundant oxygen, with oxygen levels being as high as 10% of their present atmospheric level by the end of the GOE. The sudden injection of toxic oxygen into an anaerobic biosphere may have caused the extinction of many existing anaerobic spe ...
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Mass Extinction
An extinction event (also known as a mass extinction or biotic crisis) is a widespread and rapid decrease in the biodiversity on Earth. Such an event is identified by a sharp change in the diversity and abundance of multicellular organisms. It occurs when the rate of extinction increases with respect to the background extinction rate and the rate of speciation. Estimates of the number of major mass extinctions in the last 540 million years range from as few as five to more than twenty. These differences stem from disagreement as to what constitutes a "major" extinction event, and the data chosen to measure past diversity. The "Big Five" mass extinctions In a landmark paper published in 1982, Jack Sepkoski and David M. Raup identified five particular geological intervals with excessive diversity loss. They were originally identified as outliers on a general trend of decreasing extinction rates during the Phanerozoic, but as more stringent statistical tests have been applied ...
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Retroposon
Retroposons are repetitive DNA fragments which are inserted into chromosomes after they had been reverse transcribed from any RNA molecule. Difference between retroposons and retrotransposons In contrast to retrotransposons, retroposons never encode reverse transcriptase (RT) (but see below). Therefore, they are non-autonomous elements with regard to transposition activity (as opposed to transposons). Non-long terminal repeat (LTR) retrotransposons such as the human LINE1 elements are sometimes falsely referred to as retroposons. However, this depends on the author. For example, Howard Temin published the following definition: Retroposons encode RT but are devoid of long terminal repeats (LTRs), for example long interspersed elements (LINEs). Retrotransposons also feature LTRs and retroviruses, in addition, are packaged as viral particles (virions). Retrosequences are non-autonomous elements devoid of RT. They are retroposed with the aid of the machinery of autonomous element ...
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Gas Bladder
The swim bladder, gas bladder, fish maw, or air bladder is an internal gas-filled organ that contributes to the ability of many bony fish (but not cartilaginous fish) to control their buoyancy, and thus to stay at their current water depth without having to expend energy in swimming. Also, the dorsal position of the swim bladder means the center of mass is below the center of volume, allowing it to act as a stabilizing agent. Additionally, the swim bladder functions as a resonating chamber, to produce or receive sound. The swim bladder is evolutionarily homologous to the lungs. Charles Darwin remarked upon this in ''On the Origin of Species''.Darwin, Charles (1859''Origin of Species''Page 190, reprinted 1872 by D. Appleton. Darwin reasoned that the lung in air-breathing vertebrates had derived from a more primitive swim bladder. In the embryonic stages, some species, such as redlip blenny, have lost the swim bladder again, mostly bottom dwellers like the weather fish. ...
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