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Condylarthra
Condylarthra is an informal group – previously considered an order – of extinct placental mammals, known primarily from the Paleocene and Eocene epochs. They are considered early, primitive ungulates and is now largely considered to be a wastebasket taxon, having served as a dumping ground for classifying ungulates which had not been clearly established as part of either Perissodactyla or Artiodactyla, being composed thus of several unrelated lineages. Taxonomic history Condylarthra always was a problematic group. When first described by , Phenacodontidae was the type and only family therein. , however, raised Condylarthra to an order and included a wide range of diverse placentals with generalized dentitions and postcranial skeletons. More recent researchers (i.e. post-WW2) have been more restrictive; either including only a limited number of taxa, or proposing that the term should be abandoned altogether. Due to their primitive characteristics condylarths have been consi ...
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Phenacodontidae
Phenacodontidae is an extinct family of large herbivorous mammals traditionally placed in the “wastebasket taxon” Condylarthra, which may instead represent early-stage perissodactyls. They lived from the late early Paleocene to early middle Eocene (about 60–50 million years ago) and their fossil remains have been found in North America and Europe. The only unequivocal Asian phenacodontid is '' Lophocion asiaticus''. Description These animals had a variety of body sizes, and could be as small as domestic cats ('' Tetraclaenodon'' and '' Ectocion'') and as large as sheep ('' Phenacodus''). The skull of phenacodontids is long and narrow, and equipped with a small braincase. The skeleton of phenacodontids show several primitive characteristics (the long and heavy tail for example) but also a number of advanced, Perissodactyla-like adaptations: Their long legs, for example, had five fingers, but the first finger showed a clear reduction, and in some forms (like ''Phenacod ...
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Arctocyon DB
''Arctocyon'' (from Greek ''arktos'' and ''kyôn'', "bear/dog-like") is an extinct genus of large placental mammals, part of the possibly polyphyletic family Arctocyonidae. The type species is ''A. primaevus'', though up to five other species may be known. Fossils of ''Arctocyon'' have been found in Europe and North America. ''Arctocyon'' was originally named as a subgenus of the bear-dog ''Amphicyon'', though was subsequently found to belong to a genus and family of its own. The relationship between arctocyonids and other placentals is unclear, with early classification efforts placing them as carnivores or creodonts, and later ones classifying them under Condylarthra, a wastebasket taxon for various early ungulates. More recent phylogenetic analyses suggest that arctocyonids are an artificial assemblage of several distantly-related placental lineages, and ''Arctocyon'' may be closely related to '' Loxolophus'' (another arctocyonid) and to pantodonts. ''Arctocyon'' was among the ...
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Paleocene
The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name is a combination of the Ancient Greek ''palaiós'' meaning "old" and the Eocene Epoch (which succeeds the Paleocene), translating to "the old part of the Eocene". The epoch is bracketed by two major events in Earth's history. The K–Pg extinction event, brought on by an asteroid impact (Chicxulub impact) and possibly volcanism (Deccan Traps), marked the beginning of the Paleocene and killed off 75% of species, most famously the non-avian dinosaurs. The end of the epoch was marked by the Paleocene–Eocene Thermal Maximum (PETM), which was a major climatic event wherein about 2,500–4,500 gigatons of carbon were released into the atmosphere and ocean systems, causing a spike in global temperatures and ocean acidification. ...
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Wastebasket Taxon
Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon) is a term used by some taxonomists to refer to a taxon that has the purpose of classifying organisms that do not fit anywhere else. They are typically defined by either their designated members' often superficial similarity to each other, or their ''lack'' of one or more distinct character states or by their ''not'' belonging to one or more other taxa. Wastebasket taxa are by definition either paraphyletic or polyphyletic, and are therefore not considered valid taxa under strict cladistic rules of taxonomy. The name of a wastebasket taxon may in some cases be retained as the designation of an evolutionary grade, however. Examples There are many examples of paraphyletic groups, but true "wastebasket" taxa are those that are known not to, and perhaps not intended to, represent natural groups, but are nevertheless used as convenient groups of organisms. The acritarchs are perhaps the most famous ex ...
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Astrapotheria
Astrapotheria is an extinct order of South American and Antarctic hoofed mammals that existed from the late Paleocene to the Middle Miocene, ."The uruguaytheriine Astrapotheriidae from the rich middle Miocene Honda Group of the upper Magdalena River valley in Colombia (...) are the youngest securely dated remains of that order in South America." Astrapotheres were large, rhinoceros-like animals and have been called one of the most bizarre orders of mammals with an enigmatic evolutionary history. The taxonomy of this order is not clear, but it may belong to Meridiungulata (along with Notoungulata, Litopterna, Pyrotheria and Xenungulata). In turn, Meridungulata is believed to belong to the extant superorder Laurasiatheria. Some scientists have regarded the astrapotheres (and sometimes the Meridiungulata as a whole) as members of the clade Atlantogenata. However, collagen and mitochondrial DNA sequence data analysed in 2015 places at least the notoungulates and litopterns firmly w ...
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Stapedial Branch Of Posterior Auricular Artery
In human anatomy, the stapedial branch of posterior auricular artery, or stapedial artery for short, is a small artery supplying the stapedius muscle in the inner ear. Structure In humans In humans, the stapedial artery is normally present in the fetus where it connects what is to become the external and internal carotid arteries. Part of the carotid artery system, it originates from the dorsal branch of aortic arch. Its superior supraorbital branch becomes the middle meningeal artery, while its infraorbital and mandibular branches fuses with the external carotid artery and later become the internal maxillary artery. Its trunk atrophies and is replaced by branches from the external carotid artery. In rare cases, the embryonic structure is still present after birth in which case it is referred to as a persistent stapedial artery (PSA). While the prevalence of this anomaly is unknown, it has been estimated to be present in 1 of 5,000 people. In other mammals Structures homol ...
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Petrosal Bone
The petrous part of the temporal bone is pyramid-shaped and is wedged in at the base of the skull between the sphenoid and occipital bones. Directed medially, forward, and a little upward, it presents a base, an apex, three surfaces, and three angles, and houses in its interior the components of the inner ear. The petrous portion is among the most basal elements of the skull and forms part of the endocranium. Petrous comes from the Latin word ''petrosus'', meaning "stone-like, hard". It is one of the densest bones in the body. In other mammals, it is a separate bone, the petrosal bone. The petrous bone is important for studies of ancient DNA from skeletal remains, as it tends to contain extremely well-preserved DNA. Base The base is fused with the internal surfaces of the squamous, tympanic, and mastoid parts. Apex The apex, which is rough and uneven, is received into the angular interval between the posterior border of the great wing of the sphenoid bone and the basilar ...
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Mastoid Foramen
The mastoid foramen is a hole in the posterior border of the temporal bone. It transmits an emissary vein between the sigmoid sinus and the suboccipital venous plexus, and a small branch of the occipital artery, the posterior meningeal artery to the dura mater. Structure The mastoid foramen is a hole in the posterior border of the temporal bone of the skull. The opening of the mastoid foramen is an average of 18 mm from the asterion, and around 34 mm from the external auditory meatus. It is typically very narrow. This may be around 2 mm. Variation The position and size of this foramen are very variable. It is not always present. Sometimes, it is duplicated on one side or both sides. Sometimes, it is situated in the occipital bone, or in the suture between the temporal bone and the occipital bone. Function The mastoid foramen transmits: * an emissary vein between the sigmoid sinus and the suboccipital venous plexus or the posterior auricular vein. * a small branch of ...
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Auditory Bulla
The tympanic part of the temporal bone is a curved plate of bone lying below the squamous part of the temporal bone, in front of the mastoid process, and surrounding the external part of the ear canal. It originates as a separate bone (tympanic bone), which in some mammals stays separate through life. Evolutionarily, a portion of it is derived from the angular bone of the reptilian lower jaw. Surfaces Its postero-superior surface is concave, and forms the anterior wall, the floor, and part of the posterior wall of the bony ear canal. Medially, it presents a narrow furrow, the ''tympanic sulcus'', for the attachment of the tympanic membrane. Its antero-inferior surface is quadrilateral and slightly concave; it constitutes the posterior boundary of the mandibular fossa, and is in contact with the retromandibular part of the parotid gland. Borders Its lateral border is free and rough, and gives attachment to the cartilaginous part of the ear canal. Internally, the tympanic pa ...
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Bunodont
The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone tooth", from ''mola'', millstone and ''dens'', tooth. Molars show a great deal of diversity in size and shape across the mammal groups. The third molar of humans is sometimes vestigial. Human anatomy In humans, the molar teeth have either four or five cusps. Adult humans have 12 molars, in four groups of three at the back of the mouth. The third, rearmost molar in each group is called a wisdom tooth. It is the last tooth to appear, breaking through the front of the gum at about the age of 20, although this varies among individuals and populations, and in many cases the tooth is missing. The human mouth contains upper (maxillary) and lower (mandibular) molars. They are: maxillary first molar, maxillary second molar, maxillary third molar, man ...
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Ectotympanic
The ectotympanic, or tympanicum, is a bony structure found in all mammals, located on the tympanic part of the temporal bone, which holds the tympanic membrane (eardrum) in place. In catarrhine primates (including humans), it takes a tube-shape. Its position and attachment to the skull vary between primates, and can be either inside or outside the auditory bulla. It is homologous with the angular bone of non-mammalian tetrapods A tetrapod (; from Ancient Greek τετρα- ''(tetra-)'' 'four' and πούς ''(poús)'' 'foot') is any four- limbed vertebrate animal of the clade Tetrapoda (). Tetrapods include all extant and extinct amphibians and amniotes, with the lat .... When the latter is present, it contacts the entotympanic. References External links webref: Anthropology * * Skull bones Mammal anatomy {{Vertebrate anatomy-stub ...
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Litopterna
Litopterna (from "smooth heel") is an extinction, extinct order of South American native ungulates that lived from the Paleocene to the Pleistocene-Holocene around 62.5 million to 12,000 years ago (or possibly as late as 3,500 years ago), and were also present in Antarctica during the Eocene. They represent the second most diverse group of South American ungulates after Notoungulata. It is divided into nine families, with Proterotheriidae and Macraucheniidae being the most diverse and last surviving families. Diversity The body forms of many litopterns, notably in the limb and skull structure, are broadly similar to those of living ungulates, unlike other South American native ungulate groups, which are often strongly divergent from living ungulates. Paleocene and Eocene litopterns generally had small body masses, with ''Protolipterna'' (Protolipternidae) estimated to have had a body mass of , though the Eocene Sparnotheriodontidae, sparnotheriodontids were considerably large ...
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