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Chordates
A chordate ( ) is a bilaterian animal belonging to the phylum Chordata ( ). All chordates possess, at some point during their larval or adult stages, five distinctive physical characteristics ( synapomorphies) that distinguish them from other taxa. These five synapomorphies are a notochord, a hollow dorsal nerve cord, an endostyle or thyroid, pharyngeal slits, and a post- anal tail. In addition to the morphological characteristics used to define chordates, analysis of genome sequences has identified two conserved signature indels (CSIs) in their proteins: cyclophilin-like protein and inner mitochondrial membrane protease ATP23, which are exclusively shared by all vertebrates, tunicates and cephalochordates. These CSIs provide molecular means to reliably distinguish chordates from all other animals. Chordates are divided into three subphyla: Vertebrata (fish, amphibians, reptiles, birds and mammals), whose notochords are replaced by a cartilaginous/ bony axial ...
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Yunnanozoon
''Yunnanozoon lividum'' (Yunnan + Greek ζῷον ''zôion'' (animal), with species name Latin ''lividum''; (lead-coloured), referring to preserved colour of specimens) is an extinct species of bilaterian animal from the Lower Cambrian Chengjiang biota of Yunnan province, China. Its affinities have been long the subject of controversy. Description The body of ''Yunnanozoon'' was fusiform, with specimens ranging from in length. The body was strongly laterally compressed, meaning that it was taller than it was wide. A segmented dorsal unit was present on the top of the body. The first segment of which was triangular, while the other segments were approximately rectangular. Axial stripes also ran down the body in the region immediately below the dorsal unit. Towards the front of the animal were 7 pairs of filamentous arches. These arches were covered by sac-like structures which had openings between them. Towards the back of the body a tube-like structure was present, possibly ...
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Vertebrate
Vertebrates () are animals with a vertebral column (backbone or spine), and a cranium, or skull. The vertebral column surrounds and protects the spinal cord, while the cranium protects the brain. The vertebrates make up the subphylum Vertebrata with some 65,000 species, by far the largest ranked grouping in the phylum Chordata. The vertebrates include mammals, birds, amphibians, and various classes of fish and reptiles. The fish include the jawless Agnatha, and the jawed Gnathostomata. The jawed fish include both the Chondrichthyes, cartilaginous fish and the Osteichthyes, bony fish. Bony fish include the Sarcopterygii, lobe-finned fish, which gave rise to the tetrapods, the animals with four limbs. Despite their success, vertebrates still only make up less than five percent of all described animal species. The first vertebrates appeared in the Cambrian explosion some 518 million years ago. Jawed vertebrates evolved in the Ordovician, followed by bony fishes in the Devonian. T ...
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Olfactores
Olfactores is a clade within the Chordata that comprises the Tunicata ( Urochordata) and the Vertebrata (sometimes referred to as Craniata). Olfactores represent the overwhelming majority of the phylum Chordata, as the Cephalochordata are the only chordates not included in the clade. This clade is defined by a more advanced olfactory system which, in the immediate vertebrate generation, gave rise to nostrils. Etymology The name Olfactores comes from Latin *''olfactores'' ("smellers," from purposive supine ''olfactum'' of ''olfacio'', "to smell," with plural masculine agentive nominalizing suffix ''-tores''), due to the development of pharyngeal respiratory and sensory functions, in contrast with cephalochordates such as the lancelet which lack a respiratory system and specialized sense organs. Olfactores hypothesis The long-standing Euchordata hypothesis that Cephalochordata is a sister taxon to Craniata was once widely accepted, likely influenced by significant tunicate ...
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Vetulicolia
Vetulicolia is a group of bilaterian marine animals encompassing several extinct species from the Cambrian, and possibly Ediacaran, periods. As of 2023, the majority of workers favor placing Vetulicolians in the stem group of the Chordata, but some continue to favor a more crownward placement as a sister group to the Tunicata. It was initially erected as a monophyletic clade with the rank of phylum in 2001, with subsequent work supporting its monophyly. However, more recent research suggests that vetulicolians may be paraphyletic and form a basal evolutionary grade of stem chordates. Etymology The taxon name, Vetulicolia, is derived from the type genus, '' Vetulicola'', which is a compound Latin word composed of ''vetuli'' "old" and ''cola'' "inhabitant". It was named after '' Vetulicola cuneata'', the first species of the group described in 1987. Description The vetulicolian body plan comprises two parts: a voluminous rostral (anterior) forebody, tipped with an anteri ...
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Phylum
In biology, a phylum (; : phyla) is a level of classification, or taxonomic rank, that is below Kingdom (biology), kingdom and above Class (biology), class. Traditionally, in botany the term division (taxonomy), division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about eight phyla. Current research in phylogenetics is uncovering the relationships among phyla within larger clades like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguishe ...
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Pikaia
''Pikaia gracilens'' is an extinct, primitive chordate marine animal known from the Middle Cambrian Burgess Shale of British Columbia. Described in 1911 by Charles Doolittle Walcott as an annelid, and in 1979 by Harry B. Whittington and Simon Conway Morris as a chordate, it became "the most famous early chordate fossil", or "famously known as the earliest described Cambrian chordate". It is estimated to have lived during the latter period of the Cambrian explosion. Since its initial discovery, more than a hundred specimens have been recovered. The body structure resembles that of the lancelet and it swam perhaps much like an eel. A notochord and myomeres (segmented blocks of skeletal muscles) span the entire length of the body, and are considered the defining signatures of chordate characters. Its primitive nature is indicated by the body covering, a cuticle, which is characteristic of invertebrates and some protochordates. A reinterpretation in 2024 found evidence of the ...
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Cathaymyrus
''Cathaymyrus'' is a genus of Early Cambrian chordate known from the Chengjiang biota in Yunnan Province, China. Both species have a long segmented body with no distinctive head. The segments resemble v-shaped muscle blocks found in cephalochordates such as ''Amphioxus''. A long linear impression runs along the "back" of the body looking something like a chordate notochord. While some authors have suggested that the genus is a cephalochordate and closely related to living lancelets, other authors have disputed this due to the poor preservation of the head region, considering its placement within Chordata uncertain. See also * Maotianshan shales The Maotianshan Shales () are a series of Early Cambrian sedimentary deposits in the Chiungchussu Formation or Heilinpu Formation, famous for their '' Konservat Lagerstätten'', deposits known for the exceptional preservation of fossilized orga ... * '' Haikouichthys'' * '' Myllokunmingia'' References {{Taxonbar, from=Q19890829 ...
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Caudata
The Caudata are a group of amphibians containing the extant salamanders (the order Urodela) and all extinct species of amphibians more closely related to salamanders than to frogs. They are typically characterized by a superficially lizard-like appearance, with slender bodies, blunt snouts, short limbs projecting at right angles to the body, and the presence of a tail in both larvae and adults. Disagreement exists between different authorities as to the definition of the terms "Caudata" and "Urodela". Some maintain that Urodela should be restricted to the crown group, with Caudata being used for the total group. Others restrict the name Caudata to the crown group and use Urodela for the total group. The former approach seems to be most widely adopted and is used in this article. Evolution The origins and evolutionary relationships between the three main groups of amphibians ( apodans, urodeles and anurans) is a matter of debate. A 2005 molecular phylogeny, based on rDNA a ...
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Conodont
Conodonts, are an extinct group of marine jawless vertebrates belonging to the class Conodonta (from Ancient Greek κῶνος (''kōnos''), meaning " cone", and ὀδούς (''odoús''), meaning "tooth"). They are primarily known from their hard, mineralised tooth-like structures called "conodont elements" that in life were present in the oral cavity and used to process food. Rare soft tissue remains suggest that they had elongate eel-like bodies with large eyes. Conodonts were a long-lasting group with over 300 million years of existence from the Cambrian (over 500 million years ago) to the beginning of the Jurassic (around 200 million years ago). Conodont elements are highly distinctive to particular species and are widely used in biostratigraphy as indicative of particular periods of geological time. Discovery and understanding of conodonts The teeth-like fossils of the conodont were first discovered by Heinz Christian Pander and the results published in Saint Petersburg, ...
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Classification
Classification is the activity of assigning objects to some pre-existing classes or categories. This is distinct from the task of establishing the classes themselves (for example through cluster analysis). Examples include diagnostic tests, identifying spam emails and deciding whether to give someone a driving license. As well as 'category', synonyms or near-synonyms for 'class' include 'type', 'species', 'order', 'concept', 'taxon', 'group', 'identification' and 'division'. The meaning of the word 'classification' (and its synonyms) may take on one of several related meanings. It may encompass both classification and the creation of classes, as for example in 'the task of categorizing pages in Wikipedia'; this overall activity is listed under taxonomy. It may refer exclusively to the underlying scheme of classes (which otherwise may be called a taxonomy). Or it may refer to the label given to an object by the classifier. Classification is a part of many different kinds of activ ...
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Nuucichthys
''Nuucichthys'' (meaning " Núu-ci fish") is an extinct genus of stem-group vertebrates known from the Cambrian (Miaolingian series) Marjum Formation of Utah, United States. The genus contains a single species, ''N. rhynchocephalus'', known from a single individual, representing the first Cambrian vertebrate from the Great Basin region of North America. ''Nuucichthys'' provides insights into the diversity of Cambrian vertebrates, which are more rare in Laurentian deposits. Taxonomic studies recover this genus as closely related to other Laurentian stem-vertebrates such as ''Metaspriggina'' and '' Emmonaspis'' (possible early vertebrate). History and Research The ''Nuucichthys'' holotype, UMNH.IP.6084, was discovered in sediments belonging to the House Range of Utah and sent to the Natural History Museum of Utah. The specimen consists of a well preserved individual that is flattened laterally. Although the exact origin of this fossil is uncertain, it most likely comes from the ...
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Bilateria
Bilateria () is a large clade of animals characterised by bilateral symmetry during embryonic development. This means their body plans are laid around a longitudinal axis with a front (or "head") and a rear (or "tail") end, as well as a left–right–symmetrical belly ( ventral) and back ( dorsal) surface. Nearly all bilaterians maintain a bilaterally symmetrical body as adults; the most notable exception is the echinoderms, which have pentaradial symmetry as adults, but bilateral symmetry as embryos. With few exceptions, bilaterian embryos are triploblastic, having three germ layers: endoderm, mesoderm and ectoderm, and have complete digestive tracts with a separate mouth and anus. Some bilaterians lack body cavities, while others have a primary body cavity derived from the blastocoel, or a secondary cavity, the coelom. Cephalization is a characteristic feature among most bilaterians, where the sense organs and central nerve ganglia become concentrated at th ...
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