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Archaeum
Archaea ( ) is a domain of organisms. Traditionally, Archaea only included its prokaryotic members, but this has since been found to be paraphyletic, as eukaryotes are known to have evolved from archaea. Even though the domain Archaea cladistically includes eukaryotes, the term "archaea" (: archaeon , from the Greek "ἀρχαῖον", which means ancient) in English still generally refers specifically to prokaryotic members of Archaea. Archaea were initially classified as bacteria, receiving the name archaebacteria (, in the Archaebacteria kingdom), but this term has fallen out of use. Archaeal cells have unique properties separating them from Bacteria and Eukaryota. Archaea are further divided into multiple recognized phyla. Classification is difficult because most have not been isolated in a laboratory and have been detected only by their gene sequences in environmental samples. It is unknown if they can produce endospores. Archaea are often similar to bacteria in size ...
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Promethearchaeum Syntrophicum
Promethearchaeota is the sole phylum of the kingdom Promethearchaeati, the kingdom from which the eukaryotes emerged. The phylum has one valid class: Promethearchaeia. The phylum includes all members of the group previously named Deep Sea Archaeal Group, also known as Marine Benthic Group B. A phylogenetic analysis disclosed a monophyletic grouping of the Promethearchaeota with the eukaryotes. The analysis revealed several genes with cell membrane-related functions. The presence of such genes support the hypothesis of an archaeal host for the emergence of the eukaryotes; the eocyte-like scenarios. Promethearchaeota was introduced in 2015 after the identification of a candidate genome in a metagenomic analysis of a mid-oceanic sediment sample. This analysis suggests the existence of a genus of unicellular life dubbed ''Lokiarchaeum''. The sample was taken near a hydrothermal vent at a vent field known as Loki's Castle located at the bend between Mohns/Knipovich ridge in the ...
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Nanoarchaeum Equitans
''Nanoarchaeum equitans'' is a species of marine archaea that was discovered in 2002 in a hydrothermal vent off the coast of Iceland on the Kolbeinsey Ridge by Karl Stetter. It has been proposed as the first species in a new phylum, and is the only species within the genus ''Nanoarchaeum''. Strains of this microbe were also found on the Sub-polar Mid Oceanic Ridge, and in the Obsidian Pool in Yellowstone National Park. Since it grows in temperatures approaching boiling, at about , it is considered to be a thermophile. It grows best in environments with a pH of 6, and a salinity concentration of 2%. ''Nanoarchaeum'' appears to be an obligate symbiont on the archaeon '' Ignicoccus''; it must be in contact with the host organism to survive. ''Nanoarchaeum equitans'' cannot synthesize lipids but obtains them from its host. Its cells are only 400 nm in diameter, making it the smallest known living organism, and the smallest known archaeon. ''N. equitans'' genome consists of a ...
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Thermoproteati
Thermoproteati is a kingdom of archaea. Its synonym, "TACK", is an acronym for Thaumarchaeota (now Nitrososphaerota), Aigarchaeota, Crenarchaeota (now Thermoproteota), and Korarchaeota (now Thermoproteota), the first groups discovered. They are found in different environments ranging from acidophilic thermophiles to mesophiles and psychrophiles and with different types of metabolism, predominantly anaerobic and chemosynthetic. Thermoproteati is a kingdom that is sister to the Asgard branch that gave rise to the eukaryotes. It has been proposed that the Thermoproteati kingdom be classified as "Crenarchaeota" and that the traditional "Crenarchaeota" (Thermoproteota) be classified as a class called "Sulfolobia", along with the other phyla with class rank or order. After including the kingdom category into ICNP, the only validly published name of this group is kingdom Thermoproteati (Guy and Ettema 2024). Classification * Thermoproteota (formerly "Crenarchaeota"). It is ...
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Asgard (Archaea)
Promethearchaeati (Synonym (taxonomy), syn. "Asgard") is a Kingdom (biology), kingdom belonging to the Domain (biology), domain Archaea that contain Eukaryote, eukaryotic signature proteins. It appears that the eukaryotes, the Domain (biology), domain that contains the animals, plants, Fungus, fungi and Protist, protists, Eukaryogenesis, emerged within the Promethearchaeati, in a branch containing the "''Candidatus'' Heimdallarchaeota". This supports the two-domain system of classification over the three-domain system. The kingdom contains one phylum Promethearchaeota, and it contains one valid Class (biology), class Promethearchaeia. After including the kingdom category into International Code of Nomenclature of Prokaryotes, ICNP, the only validly published name of this group is kingdom Promethearchaeati. All formerly proposed "phyla" would be de-ranked to classes in this framework. Discovery and nomenclature In the summer of 2010, sediments were analysed from a gravity Core s ...
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Archaeal Richmond Mine Acidophilic Nanoorganisms
Archaeal Richmond Mine acidophilic nanoorganisms (ARMAN) were first discovered in an extremely acidic mine located in northern California ( Richmond Mine at Iron Mountain) by Brett Baker in Jill Banfield's laboratory at the University of California Berkeley. These novel groups of archaea named ARMAN-1, ARMAN-2 ('' Candidatus Micrarchaeum acidiphilum'' ARMAN-2), and ARMAN-3 were missed by previous PCR-based surveys of the mine community because the ARMANs have several mismatches with commonly used PCR primers for 16S rRNA genes. Baker ''et al.'' detected them in a later study using shotgun sequencing of the community. The three groups were originally thought to represent three unique lineages deeply branched within the Euryarchaeota, a subgroup of the Archaea. However, based on a more complete archaeal genomic tree, they were assigned to a new kingdom named Nanobdellati. The ARMAN groups now comprise deeply divergent phyla named Micrarchaeota and Parvarchaeota. Their 16S ...
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Ignicoccus
''Ignicoccus'' is a genus of hyperthermophillic Archaea living in marine hydrothermal vents. They were discovered in samples taken at the Kolbeinsey Ridge north of Iceland, as well as at the East Pacific Rise (at 9 degrees N, 104 degrees W) in 2000. Systematics According to the comparisons of 16S rRNA genes, ''Ignicoccus'' represents a new, deeply branching lineage within the family of the Desulfurococcaceae. Three species are known: ''I. islandicus'', ''I. pacificus'' and ''I. hospitalis'' strain KIN4I. Cell structure The archaea of the genus ''Ignicoccus'' have tiny coccoid cells with a diameter of about 2 μm, that exhibit a smooth surface, an outer membrane and no S-layer. They have a previously unknown cell envelope structure—a cytoplasmic membrane, a periplasmic space (with a variable width of 20 to 400 nm, containing membrane-bound vesicles), and an outer membrane (approximately 10 nm wide, resembling the outer membrane of gram-negative bacteria). The l ...
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Nanobdellati
Nanobdellati ( syn. "DPANN") is a kingdom of archaea first proposed in 2013. Many members show novel signs of horizontal gene transfer from other domains of life. They are known as nanoarchaea or ultra-small archaea due to their smaller size (nanometric) compared to other archaea. "DPANN" is an acronym formed by the initials of the first five groups discovered: Diapherotrites, Parvarchaeota, Aenigmarchaeota, Nanoarchaeota and Nanohaloarchaeota. Later Woesearchaeota and Pacearchaeota were discovered and proposed within the "DPANN" superphylum. In 2017, another phylum Altiarchaeota was placed into the "DPANN" superphylum. The monophyly of Nanobdellati is not yet considered established, due to the high mutation rate of the included phyla, which can lead to the artifact of the long branch attraction (LBA) where the lineages are grouped basally or artificially at the base of the phylogenetic tree without being related. These analyses instead suggest that Nanobdellati belongs ...
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Phylum
In biology, a phylum (; : phyla) is a level of classification, or taxonomic rank, that is below Kingdom (biology), kingdom and above Class (biology), class. Traditionally, in botany the term division (taxonomy), division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about eight phyla. Current research in phylogenetics is uncovering the relationships among phyla within larger clades like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguishe ...
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Eukaryote
The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms are eukaryotes. They constitute a major group of Outline of life forms, life forms alongside the two groups of prokaryotes: the Bacteria and the Archaea. Eukaryotes represent a small minority of the number of organisms, but given their generally much larger size, their collective global biomass is much larger than that of prokaryotes. The eukaryotes emerged within the archaeal Kingdom (biology), kingdom Asgard (Archaea), Promethearchaeati and its sole phylum Promethearchaeota. This implies that there are only Two-domain system, two domains of life, Bacteria and Archaea, with eukaryotes incorporated among the Archaea. Eukaryotes first emerged during the Paleoproterozoic, likely as Flagellated cell, flagellated cells. The leading evolutiona ...
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Bacteria
Bacteria (; : bacterium) are ubiquitous, mostly free-living organisms often consisting of one Cell (biology), biological cell. They constitute a large domain (biology), domain of Prokaryote, prokaryotic microorganisms. Typically a few micrometres in length, bacteria were among the first life forms to appear on Earth, and are present in most of its habitats. Bacteria inhabit the air, soil, water, Hot spring, acidic hot springs, radioactive waste, and the deep biosphere of Earth's crust. Bacteria play a vital role in many stages of the nutrient cycle by recycling nutrients and the nitrogen fixation, fixation of nitrogen from the Earth's atmosphere, atmosphere. The nutrient cycle includes the decomposition of cadaver, dead bodies; bacteria are responsible for the putrefaction stage in this process. In the biological communities surrounding hydrothermal vents and cold seeps, extremophile bacteria provide the nutrients needed to sustain life by converting dissolved compounds, suc ...
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Kingdom (biology)
In biology, a kingdom is the second highest taxonomic rank, just below domain. Kingdoms are divided into smaller groups called phyla (singular phylum). Traditionally, textbooks from Canada and the United States have used a system of six kingdoms (Animalia, Plantae, Fungi, Protista, Archaea/Archaebacteria, and Bacteria or Eubacteria), while textbooks in other parts of the world, such as Bangladesh, Brazil, Greece, India, Pakistan, Spain, and the United Kingdom have used five kingdoms (Animalia, Plantae, Fungi, Protista and Monera). Some recent classifications based on modern cladistics have explicitly abandoned the term ''kingdom'', noting that some traditional kingdoms are not monophyletic, meaning that they do not consist of all the descendants of a common ancestor. The terms ''flora'' (for plants), ''fauna'' (for animals), and, in the 21st century, '' funga'' (for fungi) are also used for life present in a particular region or time. Definition and associated terms Whe ...
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Organism
An organism is any life, living thing that functions as an individual. Such a definition raises more problems than it solves, not least because the concept of an individual is also difficult. Many criteria, few of them widely accepted, have been proposed to define what an organism is. Among the most common is that an organism has autonomous reproduction, Cell growth, growth, and metabolism. This would exclude viruses, despite the fact that they evolution, evolve like organisms. Other problematic cases include colonial organisms; a colony of eusocial insects is organised adaptively, and has Germ-Soma Differentiation, germ-soma specialisation, with some insects reproducing, others not, like cells in an animal's body. The body of a siphonophore, a jelly-like marine animal, is composed of organism-like zooids, but the whole structure looks and functions much like an animal such as a jellyfish, the parts collaborating to provide the functions of the colonial organism. The evolutiona ...
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