Vessel Element
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Vessel Element
A vessel element or vessel member (also called trachea or xylem vessel) is one of the cell types found in xylem, the water conducting tissue of plants. Vessel elements are typically found in angiosperms (flowering plants) but absent from most gymnosperms such as conifers. Vessel elements are the main feature distinguishing the " hardwood" of angiosperms from the " softwood" of conifers. Morphology Xylem is the tissue in vascular plants that conducts water (and substances dissolved in it) upwards in a plant. There are two kinds of cell that are involved in the actual transport: tracheids and vessel elements. Tracheids & vessel elements make the tracheary elements of xylem.Vessel elements are the building blocks of vessels, which constitute the major part of the water transporting system in those plants in which they occur. Vessels form an efficient system for transporting water (including necessary minerals) from the root to the leaves and other parts of the plant. In seco ...
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Hardwood Pores
Hardwood is wood from dicot trees. These are usually found in broad-leaved temperate and tropical forests. In temperate and boreal latitudes they are mostly deciduous, but in tropics and subtropics mostly evergreen. Hardwood (which comes from angiosperm trees) contrasts with softwood (which is from gymnosperm trees). Characteristics Hardwoods are produced by angiosperm trees that reproduce by flowers, and have broad leaves. Many species are deciduous. Those of temperate regions lose their leaves every autumn as temperatures fall and are dormant in the winter, but those of tropical regions may shed their leaves in response to seasonal or sporadic periods of drought. Hardwood from deciduous species, such as oak, normally shows annual growth rings, but these may be absent in some tropical hardwoods. Hardwoods have a more complex structure than softwoods and are often much slower growing as a result. The dominant feature separating "hardwoods" from softwoods is the presence of ...
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Gnetales
Gnetophyta () is a division of plants (alternatively considered the subclass Gnetidae or order Gnetales), grouped within the gymnosperms (which also includes conifers, cycads, and ginkgos), that consists of some 70 species across the three relict genera: ''Gnetum'' (family Gnetaceae), ''Welwitschia'' (family Welwitschiaceae), and ''Ephedra'' (family Ephedraceae). Fossilized pollen attributed to a close relative of ''Ephedra'' has been dated as far back as the Early Cretaceous. Though diverse in the Early Cretaceous, only three families, each containing a single genus, are still alive today. The primary difference between gnetophytes and other gymnosperms is the presence of vessel elements, a system of conduits that transport water within the plant, similar to those found in flowering plants. Because of this, gnetophytes were once thought to be the closest gymnosperm relatives to flowering plants, but more recent molecular studies have brought this hypothesis into question. Thou ...
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Tracheid
A tracheid is a long and tapered lignified cell in the xylem of vascular plants. It is a type of conductive cell called a tracheary element. Angiosperms use another type of tracheary element, called vessel elements, to transport water through the xylem. The main functions of tracheid cells are to transport water and inorganic salts, and to provide structural support for trees. There are often pits on the cell walls of tracheids, which allows for water flow between cells. Tracheids are dead at functional maturity and do not have a protoplast. The wood (softwood) of gymnosperms such as pines and other conifers is mainly composed of tracheids. Tracheids are also the main conductive cells in the primary xylem of ferns. The tracheid was first named after the German botanist Carl Gustav Sanio in 1863, from the German ''Tracheide''. Evolution Tracheids were the main conductive cells found in early vascular plants. In the first 140-150 million years of vascular plant evolution, tra ...
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Aquatic Fern
The order Salviniales (formerly known as the Hydropteridales and including the former Marsileales) is an order of ferns in the class Polypodiopsida. Description Salviniales are all aquatic and differ from all other ferns in being heterosporous, meaning that they produce two different types of spores (megaspores and microspores) that develop into two different types of gametophytes (female and male gametophytes, respectively), and in that their gametophytes are endosporic, meaning that they never grow outside the spore wall and cannot become larger than the spores that produced them. The megasporangia each produce a single megaspore. In being heterosporus with endosporic gametophytes they are more similar to seed plants than to other ferns. The fertile and sterile leaves are dimorphic, taking on a different shape, and leaves bear anastomosing veins. Aerenchyma is frequently present in roots, shoots, and petioles (leaf stalks). The ferns of this order vary radically in form an ...
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Regnellidium
Regnellidium is a monotypic genus of ferns of family Marsileaceae. The single living species, ''Regnellidium diphyllum'', the two-leaf water fern, is native to southeastern Brazil and adjacent regions of Argentina. It resembles its relatives from the genus '' Marsilea'', but has 2-lobed leaves (rather than 4). This fern is sometimes grown in aquaria. A fossil assigned to the species ''Regnellidium upatoiensis'' has been found in Cretaceous deposits of the eastern United States. The genus name of ''Regnellidium'' is in honour of Anders Fredrik Regnell Anders Fredrik Regnell (8 June 1807 – 12 September 1884) was a Swedish physician and botanist. He studied in Uppsala and received his medical doctorate in 1837. As a student he served as assistant to Anders Retzius in Stockholm. He served in v ... (1807–1884), a Swedish physician and botanist. It was first described and published in Ark. Bot. Vol.3 (Issue 6) on page 2 in 1904. References Other sources * Mabberley, D. J. ...
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Marsilea
''Marsilea'' is a genus of approximately 65 species of aquatic ferns of the family Marsileaceae. The name honours Italian naturalist Luigi Ferdinando Marsili (1656–1730). These small plants are of unusual appearance and do not resemble common ferns. Common names include water clover and four-leaf clover because of the long-stalked leaves have four clover-like lobes and are either present above water or submerged. The sporocarps of some Australian species are very drought-resistant, surviving up to 100 years in dry conditions. On wetting, the gelatinous interior of the sporocarp swells, splitting it and releasing a worm-like mass that carries sori, eventually leading to germination of spores and fertilization. Uses As food Sporocarps of some Australian species such as ''Marsilea drummondii'' are edible and have been eaten by Aborigines and early white settlers, who knew it under the name ngardu or nardoo. Parts of ''Marsilea drummondii'' contain an enzyme which destroys t ...
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Pteridium Aquilinum
''Pteridium aquilinum'' (bracken, brake or common bracken), also known as eagle fern, is a species of fern occurring in temperate and subtropical regions in both hemispheres. Originally native to Eurasia and North America, the extreme lightness of its spores has led to it achieving a cosmopolitan distribution. Etymology Common bracken was first described as ''Pteris aquilina'' by Carl Linnaeus, in Volume 2 of his ''Species Plantarum'' in 1753. The origin of the specific epithet derived from the Latin ''aquila'' "eagle". In the reprint of the ''Flora Suecica'' in 1755, Linnaeus explains that the name refers to the image of an eagle seen in the transverse section of the root. In spite of this, the opinion has been forwarded that the name pertains to the shape of the mature fronds appearing akin to an eagle's wing. However, medieval scholars, including Erasmus, thought the pattern of the fibres seen in a transverse section of the stipe resembled a double-headed eagle or oak tree. Ta ...
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Selaginella
''Selaginella'' is the sole genus of vascular plants in the family Selaginellaceae, the spikemosses or lesser clubmosses. This family is distinguished from Lycopodiaceae (the clubmosses) by having scale-leaves bearing a ligule and by having spores of two types. They are sometimes included in an informal paraphyletic group called the " fern allies". '' S. moellendorffii'' is an important model organism. Its genome has been sequenced by the United States Department of Energy's Joint Genome Institute. The name ''Selaginella'' was erected by Palisot de Beauvois solely for the species ''Selaginella selaginoides'', which turns out (with the closely related ''Selaginella deflexa'') to be a clade that is sister to all other ''Selaginellas'', so any definitive subdivision of the species into separate genera leaves two taxa in ''Selaginella'', with the hundreds of other species in new or resurrected genera. ''Selaginella'' occurs mostly in the tropical regions of the world, with a ha ...
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Equisetum
''Equisetum'' (; horsetail, snake grass, puzzlegrass) is the only living genus in Equisetaceae, a family of ferns, which reproduce by spores rather than seeds. ''Equisetum'' is a "living fossil", the only living genus of the entire subclass Equisetidae, which for over 100 million years was much more diverse and dominated the understorey of late Paleozoic forests. Some equisetids were large trees reaching to tall. The genus ''Calamites'' of the family Calamitaceae, for example, is abundant in coal deposits from the Carboniferous period. The pattern of spacing of nodes in horsetails, wherein those toward the apex of the shoot are increasingly close together, is said to have inspired John Napier to invent logarithms. Modern horsetails first appeared during the Jurassic period. A superficially similar but entirely unrelated flowering plant genus, mare's tail (''Hippuris''), is occasionally referred to as "horsetail", and adding to confusion, the name "mare's tail" is someti ...
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Gnetum
''Gnetum'' is a genus of gymnosperms, the sole genus in the family Gnetaceae within the Gnetophyta. They are tropical evergreen trees, shrubs and lianas. Unlike other gymnosperms, they possess vessel elements in the xylem. Some species have been proposed to have been the first plants to be insect- pollinated as their fossils occur in association with extinct pollinating scorpionflies.Ren D, Labandeira CC, Santiago-Blay JA, Rasnitsyn A, Shih CK, Bashkuev A, Logan MA, Hotton CL, Dilcher D. (2009). Probable Pollination Mode Before Angiosperms: Eurasian, Long-Proboscid Scorpionflies. Science, 326 (5954), 840-847. Molecular phylogenies based on nuclear and plastid sequences from most of the species indicate hybridization among some of the Southeast Asian species. Fossil-calibrated molecular-clocks suggest that the ''Gnetum'' lineages now found in Africa, South America and Southeast Asia are the result of ancient long-distance dispersal across seawater.Won H, Renner SS: The internal ...
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Arthur Cronquist
Arthur John Cronquist (March 19, 1919 – March 22, 1992) was an American biologist, botanist and a specialist on Compositae. He is considered one of the most influential botanists of the 20th century, largely due to his formulation of the Cronquist system as well as being the primary co-author to the Flora of the Pacific Northwest, still the most up to date flora for three northwest U.S. States to date. Two plant genera in the aster family have been named in his honor. These are ''Cronquistia'', a possible synonym of '' Carphochaete'', and ''Cronquistianthus'', which is sometimes included as a group within ''Eupatorium''. The former was applied by R.M. King and the latter by him and Harold E. Robinson. Life Arthur Cronquist was born on March 19, 1919, in San Jose, California, but he grew up outside of Portland, Oregon, as well as in Pocatello, Idaho. His parents divorced when he was young and he and his older sister were brought up by his mother, who worked for the Union ...
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