Livoniana Multidentata
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Livoniana Multidentata
''Livoniana'' is a genus of prehistoric lobe-finned fish which lived during the Devonian period (Givetian - Frasnian stages, about 374 - 391 million years ago). This species is a transitional form between fish and the earliest tetrapods, like ''Tiktaalik'', ''Ichthyostega'' and ''Acanthostega''. Before ''Livoniana'' there was ''Elginerpeton'' and ''Obruchevichthys''. Four legs developed in water, not on land, to better escape waterliving predatory creatures like ''Hyneria ''Hyneria'' is a genus of large prehistoric predatory lobe-finned fish which lived in freshwater during the Devonian period around 360 million years ago. Etymology The genus name ''Hyneria'' is a reference to the village of Hyner, Pennsylvania, ...''. There were very lush forests, and particularly swamps, where four limbs became very useful to avoid predators. References External links ''Livoniana multidentata''at Devonian Times Elpistostegalians Prehistoric lobe-finned fish genera Transitional ...
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Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began domina ...
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Tiktaalik
''Tiktaalik'' (; Inuktitut ) is a monospecific genus of extinct sarcopterygian (lobe-finned fish) from the Late Devonian Period, about 375 Mya (million years ago), having many features akin to those of tetrapods (four-legged animals). It may have grown up to in length. Unearthed in Arctic Canada, ''Tiktaalik'' is a non-tetrapod member of Osteichthyes (bony fish), complete with scales and gills – but it has a triangular, flattened head and unusual, cleaver-shaped fins. Its fins have thin ray bones for paddling like most fish, but they also have sturdy interior bones that would have allowed Tiktaalik to prop itself up in shallow water and use its limbs for support as most four-legged animals do. Those fins and other mixed characteristics mark ''Tiktaalik'' as a crucial transition fossil, a link in evolution from swimming fish to four-legged vertebrates. This and similar animals might be the common ancestors of all vertebrate terrestrial fauna: amphibians, reptiles, birds, and ...
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Prehistoric Lobe-finned Fish Genera
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. ...
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Elpistostegalians
Elpistostegalia or Panderichthyida is an order of prehistoric lobe-finned fishes which lived during the Middle Devonian to Late Devonian period (about 385 to 374 million years ago). They represent the advanced tetrapodomorph stock, the fishes more closely related to tetrapods than the osteolepiform fishes. The earliest elpistostegalians, combining fishlike and tetrapod-like characters, are sometimes called fishapods, a phrase coined for the advanced elpistostegalian ''Tiktaalik''. Through a strict cladistic view, the order includes the terrestrial tetrapods. Palaeobiology of the elpistostegalians A rise in global oxygen content allowed for the evolution of large, predatory fish that were able to exploit the shallow tidal areas and swamplands as top predators. Several groups evolved to fill these niches, the most successful were the elpistiostegalians. In such environments, they would have been challenged by periodic oxygen deficiency. In comparable modern aquatic environments li ...
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Hyneria
''Hyneria'' is a genus of large prehistoric predatory lobe-finned fish which lived in freshwater during the Devonian period around 360 million years ago. Etymology The genus name ''Hyneria'' is a reference to the village of Hyner, Pennsylvania, near where the first specimen was found. The species epiphet ''H. lindae'' is derived from the name of the wife of Keith Stewart Thomson, who described this fish. Description ''Hyneria'' was a large fish, with discovered remains estimated between in total length. Its skull had heavy, ornamented dermal bones and its lower jaw was relatively long and shallow. The teeth were stout with those of the premaxilla forming fangs upwards of . Its body was covered by cycloid scales. It had large sensory canals to aid in detection of possible prey, as the freshwater environment it inhabited likely was murky and had low visibility. Adult individuals retained the juvenile features (i.e. partially unossified skeletons), suggesting that they were likel ...
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Obruchevichthys
''Obruchevichthys'' is an extinct genus of tetrapod from Latvia during the Late Devonian. When the jawbone, the only known fossil of this creature, was uncovered in Latvia, it was mistaken as a lobe-fin fish. However, when it was analyzed, it proved to hold many similarities to '' Elginerpeton'', from Scotland. It was then declared belonging to the earliest group of tetrapod Tetrapods (; ) are four-limb (anatomy), limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant taxon, extant and extinct amphibians, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (p ...s. References External links ''Devonian Times'' article on ''Obruchevichthys gracilis'' Further reading * ''Gaining Ground: The Origin and Early Evolution of Tetrapods'' by Jennifer A. Clack * ''Origins of the Higher Groups of Tetrapods: Controversy and Consensus'' by Hans-Peter Schultze and Linda Trueb * ''Air-Breathing Fishes: Evolution, Diversity, and Ad ...
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Elginerpeton
''Elginerpeton'' is a genus of stegocephalian (stem-tetrapod), the fossils of which were recovered from Scat Craig, Morayshire in the UK, from rocks dating to the late Devonian Period (Early Famennian stage, 368 million years ago). The only known fossil has been given the name ''Elginerpeton pancheni''. ''Elginerpeton'' is known from skeletal fragments including a partial shoulder and hip, a femur, tibia (lower hind limb), and jaw fragments. The holotype is a lower jaw fragment estimated at 40 centimeters in total length. The total body is estimated to have measured about 1.5 m (5 ft) in length. Upon its description, ''Elginerpeton'' was allied with ''Obruchevichthys'' in the family Elginerpetontidae. A biomechanical analysis of stegocephalian jaws has indicated that ''Elginerpeton'' had an unusual feeding habit among tetrapod relatives. Its jaws were thin, and plotted as the most susceptible to high stresses among the sample group. However, the heavy sculpturing of ...
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Acanthostega
''Acanthostega'' (meaning "spiny roof") is an extinct genus of stem-tetrapod, among the first vertebrate animals to have recognizable limbs. It appeared in the late Devonian period (Famennian age) about 365 million years ago, and was anatomically intermediate between lobe-finned fishes and those that were fully capable of coming onto land.


Description

The ''Acanthostega'' Polydactyly in stem-tetrapods, had eight digits on each ...
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Ichthyostega
''Ichthyostega'' (from el, ἰχθῦς , 'fish' and el, στέγη , 'roof') is an extinct genus of limbed tetrapodomorphs from the Late Devonian of Greenland. It was among the earliest four-limbed vertebrates in the fossil record, and was one of the first with weight-bearing adaptations for terrestrial locomotion. ''Ichthyostega'' possessed lungs and limbs that helped it navigate through shallow water in swamps. Although ''Ichthyostega'' is often labelled a 'tetrapod' due to the possession of limbs and fingers, it evolved long before true crown group tetrapods, and could more accurately be referred to as a stegocephalian or stem tetrapod. Likewise, while undoubtedly of amphibian build and habit, it is not considered a true member of the group in the narrow sense, as the first modern amphibians (members of the group Lissamphibia) appeared in the Triassic Period. Until finds of other early stegocephalians and closely related fishes in the late 20th century, ''Ichthyostega'' ...
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Tetrapod
Tetrapods (; ) are four-limb (anatomy), limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant taxon, extant and extinct amphibians, sauropsids (reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct therapsids and all extant mammals). Tetrapods evolved from a clade of primitive semiaquatic animals known as the Tetrapodomorpha which, in turn, evolved from ancient lobe-finned fish (sarcopterygians) around 390 million years ago in the Middle Devonian, Middle Devonian period; their forms were transitional between lobe-finned fishes and true four-limbed tetrapods. Limbed vertebrates (tetrapods in the broad sense of the word) are first known from Middle Devonian trackways, and body fossils became common near the end of the Late Devonian but these were all aquatic. The first crown group, crown-tetrapods (last common ancestors of extant tetrapods capable of terrestrial locomotion) appeared by the very early Mississippian ( ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. '' Panthera leo'' (lion) and '' Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants of an ancestral taxon are grouped together (i.e. phylogenetic analysis should c ...
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Million Years Ago
The abbreviation Myr, "million years", is a unit of a quantity of (i.e. ) years, or 31.556926 teraseconds. Usage Myr (million years) is in common use in fields such as Earth science and cosmology. Myr is also used with Mya (million years ago). Together they make a reference system, one to a quantity, the other to a particular place in a year numbering system that is ''time before the present''. Myr is deprecated in geology, but in astronomy ''Myr'' is standard. Where "myr" ''is'' seen in geology it is usually "Myr" (a unit of mega-years). In astronomy it is usually "Myr" (Million years). Debate In geology a debate remains open concerning the use of ''Myr'' (duration) plus ''Ma'' (million years ago) versus using only the term ''Ma''. In either case the term '' Ma'' is used in geology literature conforming to ISO 31-1 (now ISO 80000-3) and NIST 811 recommended practices. Traditional style geology literature is written The "ago" is implied, so that any such year number "X M ...
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