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Haldanodon
''Haldanodon'' is an extinct docodont mammaliaform which lived in the Upper Jurassic (Kimmeridgian, about 145 million years ago). Its fossil remains have been found in Portugal, in the well-known fossil locality of Guimarota, which is in the Alcobaça Formation. It may have been a semi-aquatic burrowing insectivore, similar in habits to desmans and the platypus. Several specimens are known, include a partial skeleton and well-preserved skulls. Description ''Haldanodon'' was about as long as a desman, and may have had a similar ecology. The skull was low and triangular when seen from above. For many years it had the best-known skull material of any docodont, making it vital for understanding the taxonomic position of that mammaliaform group. It was the first Mesozoic mammaliaform discovered to possess turbinal plates, a complex feature of mammalian nasal cavities. However, it also retained some curiously primitive traits in common with non-mammaliaform cynodonts. The body wa ...
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Docodont
Docodonta is an order of extinct mammaliaforms that lived during the Mesozoic, from the Middle Jurassic to Early Cretaceous. They are distinguished from other early mammaliaforms by their relatively complex molar teeth, from which the order gets its name. Until recently, Docodonta were represented primarily by teeth and jaws found across former Laurasia, (modern-day North America, Europe, and Asia). However, recent discoveries in China include some exceptionally well preserved, almost complete body fossils. Description Skeletal traits Docodonts have a long and low mandible (lower jaw), formed primarily by the tooth-bearing dentary bone. The dentary connects to the cranium via a joint with the squamosal, a connection which is strengthened relative to earlier mammaliaforms. The other bones in the jaw, known as postdentary elements, are still connected to the dentary and lie within a groove (the postdentary trough) in the rear part of the dentary’s inner edge. Nevertheless ...
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Docodontidae
Docodonta is an order of extinct mammaliaforms that lived during the Mesozoic, from the Middle Jurassic to Early Cretaceous. They are distinguished from other early mammaliaforms by their relatively complex molar teeth, from which the order gets its name. Until recently, Docodonta were represented primarily by teeth and jaws found across former Laurasia, (modern-day North America, Europe, and Asia). However, recent discoveries in China include some exceptionally well preserved, almost complete body fossils. Description Skeletal traits Docodonts have a long and low mandible (lower jaw), formed primarily by the tooth-bearing dentary bone. The dentary connects to the cranium via a joint with the squamosal, a connection which is strengthened relative to earlier mammaliaforms. The other bones in the jaw, known as postdentary elements, are still connected to the dentary and lie within a groove (the postdentary trough) in the rear part of the dentary’s inner edge. Nevertheless, ...
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Castorocauda
''Castorocauda'' is an extinct, semi-aquatic, superficially otter-like genus of docodont mammaliaforms with one species, ''C. lutrasimilis''. It is part of the Yanliao Biota, found in the Daohugou Beds of Inner Mongolia, China dating to the Middle to Late Jurassic. It was part of an explosive Middle Jurassic radiation of Mammaliaformes moving into diverse habitats and niches. Its discovery in 2006, along with the discovery of other unusual mammaliaforms, disproves the previous hypothesis of Mammaliaformes remaining evolutionarily stagnant until the extinction of the non-avian dinosaurs. Weighing an estimated , ''Castorocauda'' is the largest known Jurassic mammaliaform. It is the earliest known mammaliaform with aquatic adaptations or a fur pelt. It was also adapted for digging, and its teeth are similar to those of seals and Eocene whales, collectively suggesting it behaved similarly to the modern-day platypus and river otters and ate primarily fish. It lived in a wet, sea ...
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Mammaliaformes
Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent common ancestor of Morganucodonta and the crown group mammals; the latter is the clade originating with the most recent common ancestor of extant Monotremata, Marsupialia, and Placentalia. Besides Morganucodonta and the crown group mammals, Mammaliaformes includes Docodonta and '' Hadrocodium'' as well as the Triassic '' Tikitherium'', the earliest known member of the group. Mammaliaformes is a term of phylogenetic nomenclature. In contrast, the assignment of organisms to Mammalia has traditionally been founded on traits and, on this basis, Mammalia is slightly more inclusive than Mammaliaformes. In particular, trait-based taxonomy generally includes '' Adelobasileus'' and '' Sinoconodon'' in Mammalia, though they fall outside the Mam ...
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Mammaliaforms
Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent common ancestor of Morganucodonta and the crown group mammals; the latter is the clade originating with the most recent common ancestor of extant Monotremata, Marsupialia, and Placentalia. Besides Morganucodonta and the crown group mammals, Mammaliaformes includes Docodonta and '' Hadrocodium'' as well as the Triassic '' Tikitherium'', the earliest known member of the group. Mammaliaformes is a term of phylogenetic nomenclature. In contrast, the assignment of organisms to Mammalia has traditionally been founded on traits and, on this basis, Mammalia is slightly more inclusive than Mammaliaformes. In particular, trait-based taxonomy generally includes ''Adelobasileus'' and ''Sinoconodon'' in Mammalia, though they fall outside the Mam ...
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Mammaliaform
Mammaliaformes ("mammalian forms") is a clade that contains the crown group mammals and their closest extinct relatives; the group radiated from earlier probainognathian cynodonts. It is defined as the clade originating from the most recent common ancestor of Morganucodonta and the crown group mammals; the latter is the clade originating with the most recent common ancestor of extant Monotremata, Marsupialia, and Placentalia. Besides Morganucodonta and the crown group mammals, Mammaliaformes includes Docodonta and ''Hadrocodium'' as well as the Triassic ''Tikitherium'', the earliest known member of the group. Mammaliaformes is a term of phylogenetic nomenclature. In contrast, the assignment of organisms to Mammalia has traditionally been founded on traits and, on this basis, Mammalia is slightly more inclusive than Mammaliaformes. In particular, trait-based taxonomy generally includes '' Adelobasileus'' and '' Sinoconodon'' in Mammalia, though they fall outside the Mammaliaforme ...
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Hadrocodium
''Hadrocodium wui'' is an extinct mammaliaform that lived during the Sinemurian stage of the Early Jurassic approximately in the Lufeng Formation of the Lufeng Basin in what is now the Yunnan province in south-western China (, paleocoordinates ). It is considered as the closest relative of the class Mammalia. The fossil of this mouse-like, paper-clip sized animal was discovered in 1985 but was then interpreted as a juvenile morganucodontid. ''Hadrocodium'' remained undescribed until 2001; since then its large brain and advanced ear structure have greatly influenced the interpretation of the earliest stages of mammalian evolution, as these mammalian characters could previously be traced only to some . ''Hadrocodium'' is known only from a skull long, and the body mass is estimated to have been , making it one of the smallest Mesozoic mammaliaforms (in contrast, the related ancestral mammal would have been bigger, about ). The name ''Hadrocodium'' alludes to its large cran ...
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Mammalia
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together w ...
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Alcobaça Formation
The Alcobaça Formation, previously known as the Guimarota Formation and also known as the Consolação Unit, is a geological formation in Portugal. It dates back to the Kimmeridgian stage of the Late Jurassic. It is an important source of information on the diversity of Late Jurassic mammals. Many of the fossils were collected from the now disused and flooded Camadas de Guimarota coal mine. Vertebrate paleofauna Dinosaur eggs are geographically located in Lisbon District, Portugal. Dinosaur tracks are geographically located in Leiria District, Portugal. Amphibians Ornithischians Indeterminate euornithopod remains located in Lisbon District. Indeterminate stegosaurid remains present in Lisbon District. Saurischians Indeterminate sauropod remains located in Leiria and Lisboa."16.1 Distrito do Leiria, Portugal; 3. Camadas de Alcobaça," and "16.4 Distrito do Lisboa, Portugal; 1. Camadas de Alcobaça," in Weishampel, et al. (2004). Pages 548-549. Turtles Lepido ...
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Megazostrodon
''Megazostrodon'' is an extinct genus of basal mammaliaforms belonging to the order Morganucodonta. It is approximately 200 million years old.Fur and Fangs: Mammal Origins
. Palaeobiology and Biodiversity Research Group, University of Bristol.
Two species are known: ''M. rudnerae'' from the of and , and ''M. chenali'' from the

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Morganucodon
''Morganucodon'' ("Glamorgan tooth") is an early mammaliaform genus that lived from the Late Triassic to the Middle Jurassic. It first appeared about 205 million years ago. Unlike many other early mammaliaforms, ''Morganucodon'' is well represented by abundant and well preserved (though in the vast majority of cases disarticulated) material. Most of this comes from Glamorgan in Wales (''Morganucodon watsoni''), but fossils have also been found in Yunnan Province in China (''Morganucodon oehleri'') and various parts of Europe and North America. Some closely related animals (''Megazostrodon'') are known from exquisite fossils from South Africa. The name comes from a Latinization of ''Morganuc'', the name for South Glamorgan in the Domesday Book, the county of Wales where it was discovered by Walter Georg Kühne,Walter G. Kühne, "On a Triconodont tooth of a new pattern from a Fissure-filling in South Glamorgan", ''Proceedings of the Zoological Society of London'', volume 119 (1949� ...
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Sinoconodon
''Sinoconodon'' is an extinct genus of mammaliamorphs that appears in the fossil record of the Lufeng Formation of China in the Sinemurian stage of the Early Jurassic period, about 193 million years ago. While sharing many plesiomorphic traits with other non- mammaliaform cynodonts, it possessed a special, secondarily evolved jaw joint between the dentary and the squamosal bones, which in more derived taxa would replace the primitive tetrapod one between the articular and quadrate bones. The presence of a dentary-squamosal joint is a trait historically used to define mammals. Description This animal had skull of which suggest a presacral body length of and weight about due to the similar parameters to the European hedgehog. ''Sinoconodon'' closely resembled early mammaliaforms like '' Morganucodon'', but it is regarded as more basal, differing substantially from ''Morganucodon'' in its dentition and growth habits. Like most other non-mammalian tetrapods, such as ...
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