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Formica Truncorum
''Formica truncorum'' is a species of wood ant from the genus ''Formica''. It is distributed across a variety of locations worldwide, including central Europe and Japan. Workers can range from 3.5 to 9.0mm and are uniquely characterized by small hairs covering their entire bodies. Like all other ants, ''F. truncorum'' is eusocial and demonstrates many cooperative behaviors that are unique to its order. Colonies are either monogynous, with one queen, or polygynous, with many queens, and these two types of colonies differ in many characteristics. Range and habitat ''F. truncorum'' is located in many places worldwide, such as northern Japan, the Jura Mountains and many regions from Italy to Norway. The colonies, which can be polygynous and often polydomous (where several nests are occupied by one colony), are spread out in regions of woodland borders, where the ants will make their nests in tree stumps. Morphology ''F. truncorum'' ants are characterized by a grey-brown gaster and ...
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Johan Christian Fabricius
Johan Christian Fabricius (7 January 1745 – 3 March 1808) was a Danish zoology, zoologist, specialising in "Insecta", which at that time included all arthropods: insects, arachnids, crustaceans and others. He was a student of Carl Linnaeus, and is considered one of the most important entomologists of the 18th century, having named nearly 10,000 species of animals, and established the basis for the modern insect Biological classification, classification. Biography Johan Christian Fabricius was born on 7 January 1745 at Tønder in the Duchy of Schleswig, where his father was a doctor. He studied at the gymnasium (school), gymnasium at Altona, Hamburg, Altona and entered the University of Copenhagen in 1762. Later the same year he travelled together with his friend and relative Johan Zoëga to Uppsala University, Uppsala, where he studied under Carl Linnaeus for two years. On his return, he started work on his , which was finally published in 1775. Throughout this time, he remaine ...
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Eusociality
Eusociality (from Ancient Greek, Greek εὖ ''eu'' "good" and social), the highest level of organization of sociality, is defined by the following characteristics: cooperative Offspring, brood care (including care of offspring from other individuals), overlapping generations within a colony of adults, and a division of labor into reproductive and non-reproductive groups. The division of labor creates specialized behavioral groups within an animal society which are sometimes referred to as 'castes'. Eusociality is distinguished from all other social systems because individuals of at least one caste usually lose the ability to perform at least one behavior characteristic of individuals in another caste. Eusocial colonies can be viewed as superorganisms. Eusociality exists in certain insects, crustaceans, and mammals. It is mostly observed and studied in the Hymenoptera (ants, bees, and wasps) and in Blattodea (termites). A colony has caste differences: queens and reproductive males ...
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Allele Frequency
Allele frequency, or gene frequency, is the relative frequency of an allele (variant of a gene) at a particular locus in a population, expressed as a fraction or percentage. Specifically, it is the fraction of all chromosomes in the population that carry that allele over the total population or sample size. Microevolution is the change in allele frequencies that occurs over time within a population. Given the following: # A particular locus on a chromosome and a given allele at that locus # A population of ''N'' individuals with ploidy ''n'', i.e. an individual carries ''n'' copies of each chromosome in their somatic cells (e.g. two chromosomes in the cells of diploid species) # The allele exists in ''i'' chromosomes in the population then the allele frequency is the fraction of all the occurrences ''i'' of that allele and the total number of chromosome copies across the population, ''i''/(''nN''). The allele frequency is distinct from the genotype frequency, although they a ...
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Formica Truncorum Casent0173144 Profile 1
''Formica'' is a genus of ants of the family Formicidae, commonly known as wood ants, mound ants, thatching ants, and field ants. ''Formica'' is the type genus of the Formicidae, and of the subfamily Formicinae. The type species of genus ''Formica'' is the European red wood ant ''Formica rufa''. Ants of this genus tend to be between 4 and 8 mm long. Habitat As the name wood ant implies, many ''Formica'' species live in wooded areas where no shortage of material exists with which they can thatch their mounds (often called anthills). One shade-tolerant species is '' F. lugubris''. However, sunlight is important to most ''Formica'' species, and colonies rarely survive for any considerable period in deeply shaded, dense woodland. The majority of species, especially outside the ''F. rufa'' species group, are inhabitants of more open woodlands or treeless grassland or shrubland. In North America, at least, these habitats had a long history of frequent landscape-scale fires t ...
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Haplodiploidy
Haplodiploidy is a sex-determination system in which males develop from unfertilized eggs and are haploid, and females develop from fertilized eggs and are diploid. Haplodiploidy is sometimes called arrhenotoky. Haplodiploidy determines the sex in all members of the insect orders Hymenoptera (bees, ants, and wasps) and Thysanoptera ('thrips'). The system also occurs sporadically in some spider mites, Hemiptera, Coleoptera (bark beetles), and rotifers. In this system, sex is determined by the number of sets of chromosomes an individual receives. An offspring formed from the union of a sperm and an egg develops as a female, and an unfertilized egg develops as a male. This means that the males have half the number of chromosomes that a female has, and are haploid. The haplodiploid sex-determination system has a number of peculiarities. For example, a male has no father and cannot have sons, but he has a grandfather and can have grandsons. Additionally, if a eusocial-insect ...
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Sex Ratio
The sex ratio (or gender ratio) is usually defined as the ratio of males to females in a population. As explained by Fisher's principle, for evolutionary reasons this is typically about 1:1 in species which reproduce sexually. Many species deviate from an even sex ratio, either periodically or permanently. Examples include parthenogenic species, periodically mating organisms such as aphids, some eusocial wasps, bees, ants, and termites. The human sex ratio is of particular interest to anthropologists and demographers. In human societies, sex ratios at birth may be considerably skewed by factors such as the age of mother at birth and by sex-selective abortion and infanticide. Exposure to pesticides and other environmental contaminants may be a significant contributing factor as well. As of 2014, the global sex ratio at birth is estimated at 107 boys to 100 girls (1,000 boys per 934 girls).. Types In most species, the sex ratio varies according to the age profile of the ...
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Pheromone
A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavior of the receiving individuals. There are '' alarm pheromones'', ''food trail pheromones'', '' sex pheromones'', and many others that affect behavior or physiology. Pheromones are used by many organisms, from basic unicellular prokaryotes to complex multicellular eukaryotes. Their use among insects has been particularly well documented. In addition, some vertebrates, plants and ciliates communicate by using pheromones. The ecological functions and evolution of pheromones are a major topic of research in the field of chemical ecology. Background The portmanteau word "pheromone" was coined by Peter Karlson and Martin Lüscher in 1959, based on the Greek φερω ''pheroo'' ('I carry') and ὁρμων ''hormon'' ('stimulating'). Pheromone ...
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Hamilton's Rule
Kin selection is the evolutionary strategy that favours the reproductive success of an organism's relatives, even when at a cost to the organism's own survival and reproduction. Kin altruism can look like altruistic behaviour whose evolution is driven by kin selection. Kin selection is an instance of inclusive fitness, which combines the number of offspring produced with the number an individual can ensure the production of by supporting others, such as siblings. Charles Darwin discussed the concept of kin selection in his 1859 book, ''On the Origin of Species'', where he reflected on the puzzle of sterile social insects, such as honey bees, which leave reproduction to their mothers, arguing that a selection benefit to related organisms (the same "stock") would allow the evolution of a trait that confers the benefit but destroys an individual at the same time. R.A. Fisher in 1930 and J.B.S. Haldane in 1932 set out the mathematics of kin selection, with Haldane famously joking ...
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Relatedness
The coefficient of relationship is a measure of the degree of consanguinity (or biological relationship) between two individuals. The term coefficient of relationship was defined by Sewall Wright in 1922, and was derived from his definition of the coefficient of inbreeding of 1921. The measure is most commonly used in genetics and genealogy. A coefficient of inbreeding can be calculated for an individual, and is typically one-half the coefficient of relationship between the parents. In general, the higher the level of inbreeding the closer the coefficient of relationship between the parents approaches a value of 1, expressed as a percentage, and approaches a value of 0 for individuals with arbitrarily remote common ancestors. Coefficient of relationship The coefficient of relationship ( r ) between two individuals B and C is obtained by a summation of coefficients calculated for every line by which they are connected to their identical ancestors point, common ancestors. Each su ...
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Kin Selection
Kin selection is the evolutionary strategy that favours the reproductive success of an organism's relatives, even when at a cost to the organism's own survival and reproduction. Kin altruism can look like altruistic behaviour whose evolution is driven by kin selection. Kin selection is an instance of inclusive fitness, which combines the number of offspring produced with the number an individual can ensure the production of by supporting others, such as siblings. Charles Darwin discussed the concept of kin selection in his 1859 book, ''On the Origin of Species'', where he reflected on the puzzle of sterile social insects, such as honey bees, which leave reproduction to their mothers, arguing that a selection benefit to related organisms (the same "stock") would allow the evolution of a trait that confers the benefit but destroys an individual at the same time. R.A. Fisher in 1930 and J.B.S. Haldane in 1932 set out the mathematics of kin selection, with Haldane famously jok ...
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Nuptial Flight
Nuptial flight is an important phase in the reproduction of most ant, termite, and some bee species. It is also observed in some fly species, such as ''Rhamphomyia longicauda''. During the flight, virgin queens mating, mate with males and then land to start a new colony, or, in the case of honey bees, continue the succession of an existing hived colony. The winged version of ants and termites are known as alates. Before the flight A mature ant colony seasonally produces winged virgin queens and males, called alates. Unfertilized eggs develop into males. Fertilized eggs usually develop into wingless, sterile workers, but may develop into virgin queens if the larvae receive special attention. Within a few days after they have emerged (eclosed) from the pupa case, males are "quickly converted into single-purpose sexual missiles." Young queens and males stay in their parent colony until conditions are right for the nuptial flight. The flight requires clear weather since rain is ...
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Eusociality
Eusociality (from Ancient Greek, Greek εὖ ''eu'' "good" and social), the highest level of organization of sociality, is defined by the following characteristics: cooperative Offspring, brood care (including care of offspring from other individuals), overlapping generations within a colony of adults, and a division of labor into reproductive and non-reproductive groups. The division of labor creates specialized behavioral groups within an animal society which are sometimes referred to as 'castes'. Eusociality is distinguished from all other social systems because individuals of at least one caste usually lose the ability to perform at least one behavior characteristic of individuals in another caste. Eusocial colonies can be viewed as superorganisms. Eusociality exists in certain insects, crustaceans, and mammals. It is mostly observed and studied in the Hymenoptera (ants, bees, and wasps) and in Blattodea (termites). A colony has caste differences: queens and reproductive males ...
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