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Foregut Fermentation
Foregut fermentation is a form of digestion that occurs in the foregut of some animals such as the hamster rat, langur monkey, and the hippopotamus. It has evolved independently in several groups of mammals, and also in the hoatzin, a bird species. Foregut fermentation is employed by ruminants and pseudoruminants, some rodents and some marsupials. It has also evolved in colobine monkeys and in sloth Sloths are a Neotropical realm, Neotropical group of xenarthran mammals constituting the suborder Folivora, including the extant Arboreal locomotion, arboreal tree sloths and extinct terrestrial ground sloths. Noted for their slowness of move ...s. See also * Ruminant foregut fermentation * Hindgut fermentation * Enteric fermentation References Digestive system {{Vertebrate-anatomy-stub ...
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Digestion
Digestion is the breakdown of large insoluble food compounds into small water-soluble components so that they can be absorbed into the blood plasma. In certain organisms, these smaller substances are absorbed through the small intestine into the blood stream. Digestion is a form of catabolism that is often divided into two processes based on how food is broken down: mechanical and chemical digestion. The term mechanical digestion refers to the physical breakdown of large pieces of food into smaller pieces which can subsequently be accessed by digestive enzymes. Mechanical digestion takes place in the mouth through Chewing, mastication and in the small intestine through segmentation contractions. In chemical digestion, enzymes break down food into the small compounds that the body can use. In the human digestive system, food enters the mouth and mechanical digestion of the food starts by the action of mastication (chewing), a form of mechanical digestion, and the wetting contact ...
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Foregut
The foregut in humans is the anterior part of the alimentary canal, from the distal esophagus to the first half of the duodenum, at the entrance of the bile duct. Beyond the stomach, the foregut is attached to the abdominal walls by mesentery. The foregut arises from the endoderm, developing from the folding primitive gut, and is developmentally distinct from the midgut and hindgut. Although the term “foregut” is typically used in reference to the anterior section of the primitive gut, components of the adult gut can also be described with this designation. Pain in the epigastric region, just below the intersection of the ribs, typically refers to structures in the adult foregut. Adult foregut Components * Esophagus * Respiratory tract (lower respiratory tract) * Stomach * Duodenum (up to ampulla of vater) * Liver * Gallbladder * Pancreas * Spleen – The spleen arises from the mesodermal dorsal mesentery (the foregut arises from the endoderm not mesoderm). But the sp ...
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Hamster
Hamsters are rodents (order Rodentia) belonging to the subfamily Cricetinae, which contains 19 species classified in seven genera. They have become established as popular small pets. The best-known species of hamster is the golden or Syrian hamster (''Mesocricetus auratus''), which is the type most commonly kept as a pet. Other hamster species commonly kept as pets are the three species of dwarf hamster, Campbell's dwarf hamster (''Phodopus campbelli''), the winter white dwarf hamster (''Phodopus sungorus'') and the Roborovski hamster (''Phodopus roborovskii''), and the less common Chinese hamster (''Cricetulus griseus''). Hamsters feed primarily on seeds, fruits, vegetation, and occasionally burrowing insects. In the wild, they are crepuscular: they forage during the twilight hours. In captivity, however, they are known to live a conventionally nocturnal lifestyle, waking around sundown to feed and exercise. Physically, they are stout-bodied with distinguishing features ...
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Colobinae
The Colobinae or leaf-eating monkeys are a family (biology), subfamily of the Old World monkey family that includes 61 species in 11 genus, genera, including the black-and-white colobus, the large-nosed proboscis monkey, and the gray langurs. Some classifications split the colobine monkeys into two tribes, while others split them into three groups. Both classifications put the three African genera ''Black-and-white colobus, Colobus'', ''Red colobus, Piliocolobus'', and ''Olive colobus, Procolobus'' in one group; these genera are distinct in that they have stub thumbs (Greek κολοβός ''kolobós'' = "docked"). The various Asian genera are placed into another one or two groups. Analysis of Mitochondrial DNA, mtDNA confirms the Asian species form two distinct groups, one of langurs and the other of the "odd-nosed" species, but are inconsistent as to the relationships of the gray langurs; some studies suggest that the gray langurs are not closely related to either of these groups ...
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Hippopotamus
The hippopotamus (''Hippopotamus amphibius;'' ; : hippopotamuses), often shortened to hippo (: hippos), further qualified as the common hippopotamus, Nile hippopotamus and river hippopotamus, is a large semiaquatic mammal native to sub-Saharan Africa. It is one of only two extant species in the family Hippopotamidae, the other being the pygmy hippopotamus (''Choeropsis liberiensis'' or ''Hexaprotodon liberiensis''). Its name comes from the ancient Greek for "river horse" (). After elephants and rhinoceroses, the hippopotamus is the next largest land mammal. It is also the largest extant land artiodactyl. Despite their physical resemblance to pigs and other terrestrial even-toed ungulates, the closest living relatives of the hippopotamids are cetaceans (whales, dolphins, porpoises, etc.), from which they diverged about 55 million years ago. Hippos are recognisable for their barrel-shaped torsos, wide-opening mouths with large canine tusks, nearly hairless bodies, pillar ...
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Convergent Evolution
Convergent evolution is the independent evolution of similar features in species of different periods or epochs in time. Convergent evolution creates analogous structures that have similar form or function but were not present in the last common ancestor of those groups. The cladistic term for the same phenomenon is Cladogram#Homoplasies, homoplasy. The recurrent evolution of flight is a classic example, as flying pterygota, insects, birds, pterosaurs, and bats have independently evolved the useful capacity of flight. Functionally similar features that have arisen through convergent evolution are ''analogous'', whereas ''homology (biology), homologous'' structures or traits have a common origin but can have dissimilar functions. Bird, bat, and pterosaur wings are analogous structures, but their forelimbs are homologous, sharing an ancestral state despite serving different functions. The opposite of convergence is divergent evolution, where related species evolve different trai ...
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Hoatzin
The hoatzin ( ) or hoactzin ( ) (''Opisthocomus hoazin'') is a species of tropical bird found in swamps, riparian forests, and mangroves of the Amazon and the Orinoco basins in South America. It is the only extant species in the genus ''Opisthocomus'' which is the only extant genus in the Opisthocomidae family under the order of Opisthocomiformes. Despite being the subject of intense debate by specialists, the taxonomic position of this family is still far from clear. The hoatzin is notable for its chicks having primitive claws on two of their wing digits; the species also is unique among birds, in possessing a digestive system that non-trivially supports the fermentation and the effective breaking-down of plant matter, a trait more commonly known from herbivorous ungulate-ruminant mammals and some primates. This bird is also the national bird of Guyana, where the local name for this bird is Canje pheasant. Description The hoatzin is pheasant-sized, with a total length of ...
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Ruminant
Ruminants are herbivorous grazing or browsing artiodactyls belonging to the suborder Ruminantia that are able to acquire nutrients from plant-based food by fermenting it in a specialized stomach prior to digestion, principally through microbial actions. The process, which takes place in the front part of the digestive system and therefore is called foregut fermentation, typically requires the fermented ingesta (known as cud) to be regurgitated and chewed again. The process of rechewing the cud to further break down plant matter and stimulate digestion is called rumination. The word "ruminant" comes from the Latin ''ruminare'', which means "to chew over again". The roughly 200 species of ruminants include both domestic and wild species. Ruminating mammals include cattle, all domesticated and wild bovines, goats, sheep, giraffes, deer, gazelles, and antelopes.Fowler, M.E. (2010).Medicine and Surgery of Camelids, Ames, Iowa: Wiley-Blackwell. Chapter 1 General Biology and ...
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Pseudoruminant
Pseudoruminant is a classification of animals based on their digestive tract differing from the ruminants. Hippopotamidae, Hippopotami and camels are ungulate mammals with a three-chambered stomach (ruminants have a four-chambered stomach) while Equidae, equids (horses, Asinus, asses, zebras) and rhinoceroses are monogastric herbivores. Anatomy Like ruminants, some pseudoruminants may use foregut fermentation to break down cellulose in fibrous plant species (while most others are hindgut fermentation, hindgut fermenters with a large cecum). But they have three-chambered stomachs (while others are monogastric) as opposed to ruminant stomachs which have four compartments. Species See also * Ruminant, Traditional ruminant References

{{reflist Mammal taxonomy Animal anatomy Herbivorous mammals ...
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Rodent
Rodents (from Latin , 'to gnaw') are mammals of the Order (biology), order Rodentia ( ), which are characterized by a single pair of continuously growing incisors in each of the upper and Mandible, lower jaws. About 40% of all mammal species are rodents. They are native to all major land masses except for Antarctica, and several oceanic islands, though they have subsequently been introduced to most of these land masses by human activity. Rodents are extremely diverse in their ecology and lifestyles and can be found in almost every terrestrial habitat, including human-made environments. Species can be arboreal, fossorial (burrowing), saltatorial/ricochetal (leaping on their hind legs), or semiaquatic. However, all rodents share several morphological features, including having only a single upper and lower pair of ever-growing incisors. Well-known rodents include Mouse, mice, rats, squirrels, prairie dogs, porcupines, beavers, Cavia, guinea pigs, and hamsters. Once included wi ...
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Marsupials
Marsupials are a diverse group of mammals belonging to the infraclass Marsupialia. They are natively found in Australasia, Wallacea, and the Americas. One of marsupials' unique features is their reproductive strategy: the young are born in a relatively undeveloped state and then nurtured within a pouch on their mother's abdomen. Extant marsupials encompass many species, including kangaroos, koalas, opossums, possums, Tasmanian devils, wombats, wallabies, and bandicoots. Marsupials constitute a clade stemming from the last common ancestor of extant Metatheria, which encompasses all mammals more closely related to marsupials than to placentals. The evolutionary split between placentals and marsupials occurred 125-160 million years ago, in the Middle Jurassic-Early Cretaceous period. Presently, close to 70% of the 334 extant marsupial species are concentrated on the Australian continent, including mainland Australia, Tasmania, New Guinea, and nearby islands. The remaini ...
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Colobine
The Colobinae or leaf-eating monkeys are a subfamily of the Old World monkey family that includes 61 species in 11 genera, including the black-and-white colobus, the large-nosed proboscis monkey, and the gray langurs. Some classifications split the colobine monkeys into two tribes, while others split them into three groups. Both classifications put the three African genera '' Colobus'', ''Piliocolobus'', and '' Procolobus'' in one group; these genera are distinct in that they have stub thumbs (Greek κολοβός ''kolobós'' = "docked"). The various Asian genera are placed into another one or two groups. Analysis of mtDNA confirms the Asian species form two distinct groups, one of langurs and the other of the "odd-nosed" species, but are inconsistent as to the relationships of the gray langurs; some studies suggest that the gray langurs are not closely related to either of these groups, while others place them firmly within the langur group. Characteristics Colobines are me ...
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