Eukaryota
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Eukaryota
The eukaryotes ( ) constitute the Domain (biology), domain of Eukaryota or Eukarya, organisms whose Cell (biology), cells have a membrane-bound cell nucleus, nucleus. All animals, plants, Fungus, fungi, seaweeds, and many unicellular organisms are eukaryotes. They constitute a major group of Outline of life forms, life forms alongside the two groups of prokaryotes: the Bacteria and the Archaea. Eukaryotes represent a small minority of the number of organisms, but given their generally much larger size, their collective global biomass is much larger than that of prokaryotes. The eukaryotes emerged within the archaeal Kingdom (biology), kingdom Asgard (Archaea), Promethearchaeati and its sole phylum Promethearchaeota. This implies that there are only Two-domain system, two domains of life, Bacteria and Archaea, with eukaryotes incorporated among the Archaea. Eukaryotes first emerged during the Paleoproterozoic, likely as Flagellated cell, flagellated cells. The leading evolutiona ...
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Discoba
Excavata is an obsolete, extensive and diverse Paraphyly, paraphyletic group of unicellular Eukaryote, Eukaryota. The group was first suggested by Simpson and Patterson in 1999 and the name latinized and assigned a rank by Thomas Cavalier-Smith in 2002. It contains a variety of free-living and symbiotic protists, and includes some important parasites of humans such as ''Giardia'' and ''Trichomonas''. Excavates were formerly considered to be included in the now- obsolete Protist, Protista kingdom. They were distinguished from other lineages based on electron-microscopic information about how the cells are arranged (they have a distinctive ultrastructural identity). They are considered to be a Basal_(phylogenetics), basal flagellate lineage. On the basis of phylogenomic analyses, the group was shown to contain three widely separated eukaryote groups, the discobids, metamonads, and malawimonads. A current view of the composition of the excavates is given below, indicating that th ...
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Diaphoretickes
Diaphoretickes is a major group of eukaryote, eukaryotic organisms spanning over 400,000 species. The majority of the earth's biomass that carries out photosynthesis belongs to Diaphoretickes. In older classification systems, members of the Diaphoretickes were variously placed in the Kingdom (biology), kingdoms Protozoa or Protist, Protista. Etymology The name Diaphoretickes derives (''diaforetikés'') meaning diverse, dissimilar, referring to the wide morphology (biology), morphological and cellular diversity among members of this clade. History Eukaryotes, organisms whose cells contain a cell nucleus, nucleus, have been traditionally grouped into four kingdom (biology), kingdoms: animals, plants, fungi and protists. In the late 20th century, molecular phylogenetic analyses revealed that protists are a paraphyletic assortment of many independent evolutionary lineages or clades, from which animals, fungi and plants evolved. However, the relationships between these clades re ...
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Malawimonadida
Malawimonads (order Malawimonadida) are a small group of microorganisms with a basal position in the evolutionary tree of eukaryotes, containing only three recognized species. They're considered part of a paraphyletic group known as "Excavata". Evolution It is clear that the malawimonads are a monophyletic clade at the base of Eukaryota, but there is no consensus on the specific relationships between other basal groups, such as Discoba, Metamonada, Ancyromonadida and Podiata. The sister group to Malawimonadida varies greatly between analyses. Some phylogenetic analyses find Malawimonadida as the sister group to Podiata. Other analyses recover Malawimonadida as the sister group of Discoba or Metamonada. Very few modern analyses recover the three clades, Malawimonadida, Discoba and Metamonada, as a monophyletic Excavata. Taxonomy History The malawimonads were first described as order Malawimonadida in 2003 by Thomas Cavalier-Smith. In 2013 they were also described as a cl ...
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Amoebozoa
Amoebozoa is a major Taxonomy (biology), taxonomic group containing about 2,400 described species of Amoeba, amoeboid protists, often possessing blunt, fingerlike, Pseudopod#Morphology, lobose pseudopods and tubular mitochondrial cristae. In traditional classification schemes, Amoebozoa is usually ranked as a phylum within either the kingdom (biology), kingdom Protista or the kingdom Protozoa. In the classification favored by the International Society of Protistologists, it is retained as an unranked "supergroup (biology), supergroup" within Eukaryota. Molecular genetics, Molecular genetic analysis supports Amoebozoa as a monophyletic clade. Modern studies of eukaryotic phylogenetic trees identify it as the sister group to Opisthokonta, another major clade which contains both fungi and animals as well as several other clades comprising some 300 species of unicellular eukaryotes. Amoebozoa and Opisthokonta are sometimes grouped together in a high-level taxon, named Amorphea. Amoeboz ...
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Rhizaria
The Rhizaria are a diverse and species-rich clade of mostly unicellular eukaryotes. Except for the Chlorarachniophytes and three species in the genus '' Paulinella'' in the phylum Cercozoa, they are all non-photosynthetic, but many Foraminifera and Radiolaria have a symbiotic relationship with unicellular algae. A multicellular form, ''Guttulinopsis vulgaris'', a cellular slime mold, has been described. This group was used by Cavalier-Smith in 2002, although the term "Rhizaria" had been long used for clades within the currently recognized taxon. Being described mainly from rDNA sequences, they vary considerably in form, having no clear morphological distinctive characters ( synapomorphies), but for the most part they are amoeboids with filose, reticulose, or microtubule-supported pseudopods. In the absence of an apomorphy, the group is ill-defined, and its composition has been very fluid. Some Rhizaria possess mineral exoskeletons ( thecae or loricas), which are in diffe ...
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Alveolate
The alveolates (meaning "pitted like a honeycomb") are a group of protists, considered a major unranked clade or superphylum within Eukaryota. They are currently grouped with the Stramenopiles and Rhizaria among the protists with tubulocristate mitochondria into the SAR supergroup. Characteristics The most notable shared characteristic is the presence of cortical (near the surface) alveoli (sacs). These are flattened vesicles (sacs) arranged as a layer just under the membrane and supporting it, typically contributing to a flexible pellicle (thin skin). In armored dinoflagellates they may contain stiff plates. Alveolates have mitochondria with tubular cristae ( invaginations), and cells often have pore-like intrusions through the cell surface. The group contains free-living and parasitic organisms, predatory flagellates, and photosynthetic organisms. Almost all sequenced mitochondrial genomes of ciliates and apicomplexa are linear. The mitochondria almost all carry mtDN ...
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Telonemia
Telonemia is a phylum of microscopic eukaryotes commonly known as telonemids. They are unicellular free-living flagellates with a unique combination of Cell (biology), cell structures, including a highly complex cytoskeleton unseen in other eukaryotes. Telonemia shares several distinctive features with its related group, the SAR supergroup, SAR Supergroup (biology), supergroup. Among these features are cortical alveoli, small sacs beneath the cell's surface that act as cushions, providing support and helping to maintain the cell's shape. Additionally, they possess tripartite mastigonemes, complex three-part hair-like structures on their flagella, the whip-like tails used for movement. These structures enhance their swimming capabilities by increasing resistance against water. Furthermore, Telonemia is equipped with filopodia, very thin, thread-like projections extending from the cell body. These projections can serve various purposes, such as aiding in movement or capturing foo ...
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Fungi
A fungus (: fungi , , , or ; or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and mold (fungus), molds, as well as the more familiar mushrooms. These organisms are classified as one of the kingdom (biology)#Six kingdoms (1998), traditional eukaryotic kingdoms, along with Animalia, Plantae, and either Protista or Protozoa and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of motility, mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related o ...
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Animal
Animals are multicellular, eukaryotic organisms in the Biology, biological Kingdom (biology), kingdom Animalia (). With few exceptions, animals heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, have myocytes and are motility, able to move, can reproduce sexually, and grow from a hollow sphere of Cell (biology), cells, the blastula, during embryonic development. Animals form a clade, meaning that they arose from a single common ancestor. Over 1.5 million extant taxon, living animal species have been species description, described, of which around 1.05 million are insects, over 85,000 are molluscs, and around 65,000 are vertebrates. It has been estimated there are as many as 7.77 million animal species on Earth. Animal body lengths range from to . They have complex ecologies and biological interaction, interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as ...
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Provora
Provora is a supergroup of eukaryotes made up of predatory microbes, whose name implies something such as "protists devouring voraciously". It was reported that ten strains were isolated and cultured in 2022. They are predators of other microorganisms. Their discovery was very delayed, compared to other microorganisms in their environments, due to their rarity. Their 18S ribosomal RNA is very different from that of other eukaryotes consistent with their being a lineage without known close relatives; this was confirmed by phylogenomic analyses of datasets of several hundred proteins, so they were taxonomically placed in a separate supergroup. Evolution External relationships The supergroup Provora is composed of microbes that form an ancient lineage of eukaryotes. The initial phylogenetic analyses recovered a position within the larger supergroup Diaphoretickes, particularly as sister to the protist groups TSAR and Haptista. However, a 2024 phylogenomic analysis recovered Provo ...
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Hemimastigophora
Hemimastigophora is a group of unicellular, single-celled eukaryotic organisms including the Spironematellidae, first identified in 1988, and the Paramastigidae. Over the next 30 years, different authors proposed placing these organisms in various branches of the eukaryotes. In 2018 Lax ''et al''. reported the first genetic information for Spironemidae, and suggest that they are from an ancient lineage of eukaryotes which constitute a separate clade from all other eukaryotic kingdom (biology), kingdoms. It may be related to the Telonemia. History of classification Hemimastigophora was established in 1988 by Foissner ''et al''., as a new phylum with a single family, Spironemidae. Its placement on the eukaryote Tree of life (biology), tree of life was unclear, but the authors suggested that the structure of its Pellicle (biology), pellicle and cell nucleus indicated a close relationship with Euglenozoa. For 30 years after the description of the group, no genetic informati ...
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Ancyromonadida
Ancyromonadida or Planomonadida is a small group of biflagellated eukaryotes found in the soil and in aquatic habitats, where they feed on bacteria.Cavalier-Smith, T. (2013)Early evolution of eukaryote feeding modes, cell structural diversity, and classification of the protozoan phyla Loukozoa, Sulcozoa, and Choanozoa European journal of protistology, 49(2), 115-178. They are freshwater or marine organisms, benthic, dorsoventrally compressed and with two unequal flagellae, each emerging from a separate pocket. The apical anterior flagellum can be very thin or end in the cell membrane, while the posterior flagellum is long and is inserted ventrally or laterally. The cell membrane is supported by a thin single-layered theca and the mitochondrial crests are discoidal/flat. The group's placement is doubtful, as it seems to fall outside the five supergroups of eukaryotes. Cavalier-Smith considers that they constitute a basal group to Amoebozoa and Opisthokonta and places it togeth ...
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