Decapod Anatomy
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Decapod Anatomy
The decapod (crustaceans such as a crab, lobster, shrimp or prawn) is made up of 20 body segments grouped into two main body parts: the cephalothorax and the pleon (abdomen). Each segment may possess one pair of appendages, although in various groups these may be reduced or missing. They are, from head to tail: Cephalothorax Head #antennules # antennae #mandibles # first maxillae # second maxillae The head also bears the (usually stalked) compound eyes. The distal portion of a mandible or maxilla which has a sensory function is known as a palp. Thorax / pereon #first maxillipeds #second maxillipeds #third maxillipeds #first pereiopods #second pereiopods #third pereiopods #fourth pereiopods #fifth pereiopods Maxillipeds are appendages modified to function as mouthparts. Particularly in the less advanced decapods, these can be very similar to the pereiopods. Pereiopods are primarily walking legs and are also used for gathering food. They are also the ten legs from which decapods ...
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Decapoda
The Decapoda or decapods (literally "ten-footed") are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crabs, lobsters, crayfish, shrimp and prawns. Most decapods are scavengers. The order is estimated to contain nearly 15,000 species in around 2,700 genera, with around 3,300 fossil species. Nearly half of these species are crabs, with the shrimp (about 3,000 species) and Anomura including hermit crabs, porcelain crabs, squat lobsters (about 2500 species) making up the bulk of the remainder. The earliest fossil decapod is the Devonian '' Palaeopalaemon''. Anatomy Decapods can have as many as 38 appendages, arranged in one pair per body segment. As the name Decapoda (from the Greek , ', "ten", and , '' -pod'', "foot") implies, ten of these appendages are considered legs. They are the pereiopods, found on the last five thoracic segments. In many decapods, one pair of these "legs" has enlarged pincers, called chelae, with the leg ...
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Hermit Crab
Hermit crabs are anomuran decapod crustaceans of the superfamily Paguroidea that have adapted to occupy empty scavenged mollusc shells to protect their fragile exoskeletons. There are over 800 species of hermit crab, most of which possess an asymmetric abdomen concealed by a snug-fitting shell. Hermit crabs' soft (non-calcified) abdominal exoskeleton means they must occupy shelter produced by other organisms or risk being defenseless. The strong association between hermit crabs and their shelters has significantly influenced their biology. Almost 800 species carry mobile shelters (most often calcified snail shells); this protective mobility contributes to the diversity and multitude of crustaceans found in almost all marine environments. In most species, development involves metamorphosis from symmetric, free-swimming larvae to morphologically asymmetric, benthic-dwelling, shell-seeking crabs. Such physiological and behavioral extremes facilitate a transition to a sheltered lif ...
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Euphausia Superba
Antarctic krill (''Euphausia superba'') is a species of krill found in the Antarctic waters of the Southern Ocean. It is a small, swimming crustacean that lives in large schools, called swarms, sometimes reaching densities of 10,000–30,000 individual animals per cubic metre. It feeds directly on minute phytoplankton, thereby using the primary production energy that the phytoplankton originally derived from the sun in order to sustain their pelagic (open ocean) life cycle. It grows to a length of , weighs up to , and can live for up to six years. It is a key species in the Antarctic ecosystem and in terms of biomass, is one of the most abundant animal species on the planet (approximately 500 million tons, corresponding to 300 to 400 trillion individuals). Life cycle The main spawning season of Antarctic krill is from January to March, both above the continental shelf and also in the upper region of deep sea oceanic areas. In the typical way of all krill, the male attaches a s ...
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Carcinisation
Carcinisation (or carcinization) is an example of convergent evolution in which a crustacean evolves into a crab-like form from a non-crab-like form. The term was introduced into evolutionary biology by L. A. Borradaile, who described it as "one of the many attempts of Nature to evolve a crab". Most carcinised crustaceans belong to the infraorder Anomura. Definition of carcinised morphology It was stated by Lancelot Alexander Borradaile in 1916 that: Keiler et al., 2017 defines a carcinised morphology as follows: * "The carapace is flatter than it is broad and possesses lateral margins" * "The sternites are fused into a wide sternal plastron which possesses a distinct emargination on its posterior margin." * "The pleon is flattened and strongly bent, in dorsal view completely hiding the tergites of the fourth pleonal segment, and partially or completely covers the plastron" Examples Carcinisation is believed to have occurred independently in at least five ...
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Crab
Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all the world's oceans, in freshwater, and on land, are generally covered with a thick exoskeleton, and have a single pair of pincers. They first appeared during the Jurassic Period. Description Crabs are generally covered with a thick exoskeleton, composed primarily of highly mineralized chitin, and armed with a pair of chelae (claws). Crabs vary in size from the pea crab, a few millimeters wide, to the Japanese spider crab, with a leg span up to . Several other groups of crustaceans with similar appearances – such as king crabs and porcelain crabs – are not true crabs, but have evolved features similar to true crabs through a process known as carcinisation. Environment Crabs are found in all of the world's oceans, as well as in ...
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Caridoid Escape Reaction
The caridoid escape reaction, also known as lobstering or tail-flipping, refers to an innate escape mechanism in marine and freshwater crustaceans such as lobsters, krill, shrimp and crayfish. The reaction, most extensively researched in crayfish, allows crustaceans to escape predators through rapid abdominal flexions that produce powerful swimming strokes—thrusting the crustacean backwards through the water and away from danger. The type of response depends on the part of the crustacean stimulated, but this behavior is complex and is regulated both spatially and temporally through the interactions of several neurons. Discovery of the first command neuron-mediated behavior In 1946, C. A. G. Wiersma first described the tail-flip escape in the crayfish '' Procambarus clarkii'' and noted that the giant interneurons present in the tail were responsible for the reaction. The aforementioned neuronal fibres consist of a pair of lateral giant interneurons and a pair of medial giant i ...
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Telson
The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on account of not arising in the embryo from teloblast areas as other segments. It never carries any appendages, but a forked "tail" called the caudal furca may be present. The shape and composition of the telson differs between arthropod groups. Crustaceans In lobsters, shrimp and other decapods, the telson, along with the uropods, forms the tail fan. This is used as a paddle in the caridoid escape reaction ("lobstering"), whereby an alarmed animal rapidly flexes its tail, causing it to dart backwards. Krill can reach speeds of over 60 cm per second by this means. The trigger time to optical stimulus is, in spite of the low temperatures, only 55 milliseconds. In the Isopoda and Tanaidacea (superorder Peracarida), the last abdom ...
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Uropod
Uropods are posterior appendages found on a wide variety of crustaceans. They typically have functions in locomotion. Definition Uropods are often defined as the appendages of the last body segment of a crustacean. An alternative definition suggested by Frederick R. Schram restricts the term to those structures arising from the segment before the anal segment (the segment which carries the anus). Under this latter definition, the appendages of the anal segment are caudal rami, which are analogous to uropods. Form Uropods are typically biramous – comprising an endopod and an exopod. The exopod is typically the larger, and may be divided in two by a transverse suture known as the diaeresis. The uropods may work in concert with the telson The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment o ...
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Biramous
The arthropod leg is a form of jointed appendage of arthropods, usually used for walking. Many of the terms used for arthropod leg segments (called podomeres) are of Latin origin, and may be confused with terms for bones: ''coxa'' (meaning hip, plural ''coxae''), ''trochanter'', ''femur'' (plural ''femora''), ''tibia'' (plural ''tibiae''), ''tarsus'' (plural ''tarsi''), ''ischium'' (plural ''ischia''), ''metatarsus'', ''carpus'', ''dactylus'' (meaning finger), ''patella'' (plural ''patellae''). Homologies of leg segments between groups are difficult to prove and are the source of much argument. Some authors posit up to eleven segments per leg for the most recent common ancestor of extant arthropods but modern arthropods have eight or fewer. It has been argued that the ancestral leg need not have been so complex, and that other events, such as successive loss of function of a ''Hox''-gene, could result in parallel gains of leg segments. In arthropods, each of the leg segments arti ...
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Gonopods
Gonopods are specialized appendages of various arthropods used in reproduction or egg-laying. In males, they facilitate the transfer of sperm from male to female during mating, and thus are a type of intromittent organ. In crustaceans and millipedes, gonopods are modified walking or swimming legs. Gonopods may be highly decorated with elaborate structures which may play roles in sperm competition, and can be used to differentiate and identify closely related species. Gonopods generally occur in one or more pairs, as opposed to the single (un-paired) reproductive organs such as the aedeagus of insects or the penis of harvestmen. Insects In insects, gonopods are appendages of the genital segment that may be used in insemination, or that comprise the egg-laying apparatus. Crustaceans In male decapod crustaceans, gonopods are modified swimming appendages ( pleopods). The anterior two pair of pleopods in males are modified for sperm transferring, with differing degree of morph ...
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Dendrobranchiata
Dendrobranchiata is a suborder of decapods, commonly known as prawns. There are 540 extant species in seven families, and a fossil record extending back to the Devonian. They differ from related animals, such as Caridea and Stenopodidea, by the branching form of the gills and by the fact that they do not brood their eggs, but release them directly into the water. They may reach a length of over and a mass of , and are widely fished and farmed for human consumption. Shrimp and prawns While Dendrobranchiata and Caridea belong to different suborders of Decapoda, they are very similar in appearance, and in many contexts such as commercial farming and fisheries, they are both often referred to as "shrimp" and "prawn" interchangeably. In the United Kingdom, the word "prawn" is more common on menus than "shrimp", while the opposite is the case in North America. The term "prawn" is also loosely used to describe any large shrimp, especially those that come 15 (or fewer) to the poun ...
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Uropods
Uropods are posterior appendages found on a wide variety of crustaceans. They typically have functions in locomotion. Definition Uropods are often defined as the appendages of the last body segment of a crustacean. An alternative definition suggested by Frederick R. Schram restricts the term to those structures arising from the segment before the anal segment (the segment which carries the anus). Under this latter definition, the appendages of the anal segment are caudal rami, which are analogous to uropods. Form Uropods are typically biramous – comprising an endopod and an exopod. The exopod is typically the larger, and may be divided in two by a transverse suture known as the diaeresis. The uropods may work in concert with the telson The telson () is the posterior-most division of the body of an arthropod. Depending on the definition, the telson is either considered to be the final segment of the arthropod body, or an additional division that is not a true segment on ...
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