Smilacaceae
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Smilacaceae
Smilacaceae, the greenbriers, is a family of flowering plants. While they were often assigned to a more broadly defined family Liliaceae, most recent botanists have accepted the two as distinct families, diverging around 55 million years ago during the Early Paleogene. One characteristic that distinguishes Smilacaceae from most of the other members of the Liliaceae-like Liliales is that it has true vessels in its conducting tissue. Another is that the veins of the leaves, between major veins, are reticulate (net-shaped), rather than parallel as in most monocots. Taxonomy The APG II system, of 2003 (unchanged from the APG system, of 1998), recognizes this family and places it in the order Liliales, in the clade monocots. Earlier it was a family of two genera, '' Heterosmilax'' and ''Smilax'', but DNA studies have shown that ''Heterosmilax'' has arisen from ''Smilax'' and the two genera are now merged. This results in Smilax being the only genus in Smilacaceae with about 210 kno ...
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Liliales
Liliales is an order (biology), order of monocotyledonous flowering plants in the Angiosperm Phylogeny Group and Angiosperm Phylogeny Web List of systems of plant classification, system, within the lilioid monocots. This order of necessity includes the family (biology), family Liliaceae. The APG III system (2009) places this order in the monocot clade. In APG III, the family Luzuriagaceae is combined with the family Alstroemeriaceae and the family Petermanniaceae is recognized. Both the order Lililiales and the family Liliaceae have had a widely disputed history, with the circumscription (taxonomy), circumscription varying greatly from one taxonomist to another. Previous members of this order, which at one stage included most monocots with conspicuous tepals and lacking starch in the endosperm are now distributed over three orders, Liliales, Dioscoreales and Asparagales, using predominantly molecular phylogenetics. The newly delimited Liliales is monophyletic, with ten families. W ...
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