Scarabaeiformia
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Scarabaeiformia
Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Its constituent families are also undergoing revision presently, and the family list below is only preliminary. The oldest confirmed member of the group is '' Alloioscarabaeus'' from the Middle Jurassic Jiulongshan Formation of Inner Mongolia, China. Families The following families are listed in Bouchard (2011): * Belohinidae Paulian, 1959 * Diphyllostomatidae Holloway, 1972 ( false stag beetles) * Geotrupidae Latreille, 1802 ( earth-boring dung beetles) * Glaphyridae MacLeay, 1819 (bumble bee scarab beetles) * Glaresidae Kolbe, 1905 (enigmatic scarab beetles) * Hybosoridae Erichson, 1847 ( scavenging scarab beetles) ** inclusive of Ceratocanthidae (pill scarab beetles) * Lucanidae Latreille 1804 (stag beetles) * Ochodaeidae Mulsant and Rey 1871 (sand-loving scarab beetles) ...
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Ochodaeidae
Ochodaeidae, also known as the sand-loving scarab beetles, is a small family of scarabaeiform beetles occurring in many parts of the world. These beetles are small, ranging from . Their bodies are elongate and convex, with black and brown colors including yellowish- and reddish-brown shades. As of 2012, the biology and habits of Ochodaeidae beetles is still mostly unknown. Most types have been collected in sandy areas at night, while some of their species are active during the day. Taxonomy Ochodaeidae beetles belong to the infraorder Scarabaeiformia, which contains only one superfamily, the Scarabaeoidea. The most striking feature of the Scarabaeoidea are the ends of their antennae, that are divided into several lamellae, thus resembling a fan. Another distinguishing feature are their legs, that possess teeth and are adapted for digging. Ochodaeidae is divided into two subfamilies containing five tribes and 15 genera Genus ( plural genera ) is a taxonomic rank used in ...
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Hybosoridae
Hybosoridae, sometimes known as the scavenger scarab beetles, is a family of scarabaeiform beetles. The 690 species in 97 genera occur widely in the tropics, but little is known of their biology. Hybosorids are small, 5–7 mm in length and oval in shape. Color ranges from a glossy light brown to black. They are distinctive for their large mandibles and labrum, and their 10-segmented antennae, in which the 8th antennomore of the club is deeply grooved and occupied by the 9th and 10th antennomeres. The legs have prominent spurs. The larvae have the C-shape and creamy white appearance typical of the scarabaeiforms. The 4-segmented legs are well-developed; the front legs are used to stridulate by rubbing against the margin of the epipharynx, a habit unique to this family. Adults are known to feed on invertebrate and vertebrate carrion, with some found in dung. Larvae have been found in decomposing plant material. Little more is known of their life histories. The group has ...
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Diphyllostomatidae
The false stag beetles (''Diphyllostoma'') are a group of three species of rare beetles known only from California. Almost nothing is known of their life history beyond that the adults are diurnal and females are flightless; larvae have not been observed. Their length ranges from 5 to 9 mm; bodies are elongate, with a generally dull brown to reddish-brown color. Both body and legs are covered with longish hairs. Originally classed with the Lucanidae, ''Diphyllostoma'' have a number of characteristics not shared with any other type of stag beetle, and so in 1972 Holloway proposed a separate family Diphyllostomatidae, which has since been accepted. Species * '' Diphyllostoma fimbriatum'' (Fall), 1901 * '' Diphyllostoma linsleyi'' Fall, 1932 * '' Diphyllostoma nigricolle'' (Fall), 1901 Notes References * Mary Liz Jameson and Brett C. Ratcliffe, "Diphyllostomatidae", in Ross H. Arnett, Jr. Ross Harold Arnett Jr. (April 13, 1919 – July 16, 1999) was an American en ...
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Ceratocanthidae
''Ceratocanthinae'' is a subfamily of the scarabaeoid beetle family Hybosoridae Hybosoridae, sometimes known as the scavenger scarab beetles, is a family of scarabaeiform beetles. The 690 species in 97 genera occur widely in the tropics, but little is known of their biology. Hybosorids are small, 5–7 mm in length a .... It includes three tribes comprising 43 genera and 366 species; it was formerly treated as a separate family, Ceratocanthidae. Description Ceratocanthinae are small sized beetles from 2.0 to 10.0 millimeters in length. Adult beetles can be found on the bark and branches of dead trees and on fungus. Distribution Ceratocanthinae are relatively widespread. They can be found in Australian, Afrotropical, Indomalaysian, Neotropical, Nearctic, and Palaearctic regions. Ecology The adults have been found to Association (ecology), associate with termites and ants. Larvae live under bark and in burrows of bessbugs (Passalidae). Taxonomy The subfamily Ceratoc ...
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Glaresidae
''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, the Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy areas worldwide (except Australia), only the nocturnal adults have ever been collected (typically at lights), and both the larvae and biology of ''Glaresis'' are as yet unknown. Due to their narrow habitat associations, a great number of these species occur in extremely limited geographic areas, and are accordingly imperiled by habitat destruction. These beetles are small, 2.5–6 mm long, and have the stocky appearance typical of fossorial scarabs, with short, heavy, spurred legs. Color ranges from tan to dark brown, and the back is covered with short setae. Efforts to raise glaresids in the laboratory were undertaken in the 1980s by C. H. Scholtz and others, but were unsuccessful. ''Glaresis'' was originally classified with Trogidae ( ...
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Rain Beetle
The rain beetles are a group of beetles whose extant species are found only in the far west of North America. They spend most of their lives underground, emerging in response to rain or snow, thus the common name. Formerly classified in the Scarabaeidae (and later the Geotrupidae), they are currently assigned to their own family Pleocomidae, considered the sister group to all the remaining families of Scarabaeoidea. The family contains a single extant genus, '' Pleocoma'', and two extinct genera, ''Cretocoma'', described in 2002 from Late Cretaceous deposits in Mongolia, and '' Proteroscarabeus'' of Late Cretaceous China.Krell, Frank-Thorsten. "The fossil record of Mesozoic and Tertiary Scarabaeoidea (Coleoptera: Polyphaga)." Invertebrate Systematics 14.6 (2000): 871-905. Extant members of '' Pleocoma'' are known from extreme southern Washington, throughout the mountains of Oregon and California, and into the extreme north of Baja California Baja California (; 'Lower Califo ...
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Belohinidae
''Belohina inexpectata'' is a species of polyphaga Polyphaga is the largest and most diverse suborder of beetles. It comprises 144 families in 16 superfamilies, and displays an enormous variety of specialization and adaptation, with over 350,000 described species, or approximately 90% of the b ...n beetles and the sole member of family Belohinidae. It is endemic to southern Madagascar. Only a few specimens of this species are known. References Scarabaeoidea genera Monotypic Scarabaeiformia genera {{Scarabaeoidea-stub ...
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False Stag Beetle
The false stag beetles (''Diphyllostoma'') are a group of three species of rare beetles known only from California. Almost nothing is known of their life history beyond that the adults are diurnal and females are flightless; larvae have not been observed. Their length ranges from 5 to 9 mm; bodies are elongate, with a generally dull brown to reddish-brown color. Both body and legs are covered with longish hairs. Originally classed with the Lucanidae, ''Diphyllostoma'' have a number of characteristics not shared with any other type of stag beetle, and so in 1972 Holloway proposed a separate family Diphyllostomatidae, which has since been accepted. Species * ''Diphyllostoma fimbriatum'' (Fall), 1901 * ''Diphyllostoma linsleyi'' Fall, 1932 * ''Diphyllostoma nigricolle'' (Fall), 1901 Notes References * Mary Liz Jameson and Brett C. Ratcliffe, "Diphyllostomatidae", in Ross H. Arnett, Jr. and Michael C. Thomas, ''American Beetles'' (CRC Press The CRC Pres ...
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Passalidae
Passalidae is a family of beetles known variously as "bessbugs", "bess beetles", "betsy beetles" or "horned passalus beetles". Nearly all of the 500-odd species are tropical; species found in North America are notable for their size, ranging from 20–43 mm, for having a single "horn" on the head, and for a form of social behavior unusual among beetles. Bodies elongate-cylindrical and black overall; ventral surfaces may be covered with yellow setae. The head is narrower than the thorax, with antennae consisting of 10 antennomeres with a three-segment club. The elytra are elongate with parallel sides, and heavily striated. They are subsocial (brood caring) beetles that live in groups within rotting logs or stumps. The beetles will excavate tunnel systems within rotting wood where the females then lay their eggs. They care for their young by preparing food for them and helping the larvae construct the pupal case. Both adults and larvae must consume adult feces which have be ...
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