Heterostraci Genera
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Heterostraci Genera
Heterostraci (Ancient Greek, ἕτερος+ὄστρακον "those itha different shell" i is pl. of -us is an extinct subclass of pteraspidomorph, ostracoderm, jawless vertebrate that lived primarily in marine and estuary environments. Heterostraci existed from the mid-Ordovician to the conclusion of the Devonian. Description and anatomy The heterostracans differed from other Paleozoic agnathan taxa both in the arrangement and histology of their scales. Most heterostracans had two plates which form a large dorsal shield and a large ventral shield, and had series of scales arranged in various patterns on the sides of their bodies, the exact pattern differing from one group to another. In a few primitive forms, such as ''Lepidaspis'', the dorsal and ventral shields are composed of a mosaic of tiny scales. In most other known forms, though, these tiny scales have fused together to form the shield-plates. The scales of heterostracans are histologically distinct from othe ...
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Wenlock (Silurian)
The Wenlock Epoch (sometimes referred to as the Wenlockian) is the second epoch of the Silurian. It is preceded by the Llandovery Epoch and followed by the Ludlow Epoch. Radiometric dates constrain the Wenlockian between and million years ago. Naming and history The Wenlock is named after Wenlock Edge, an outcrop of rocks near the town of Much Wenlock in Shropshire ( West Midlands, United Kingdom). The name was first used in the term "Wenlock and Dudley rocks" by Roderick Murchison in 1834 to refer to the limestones and underlying shales that underlay what he termed the "Ludlow rocks". He later modified this term to simply the "Wenlock rocks" in his book, ''The Silurian System'' in 1839. Definition and subdivision The Wenlock's beginning is defined by the lower boundary (or GSSP) of the Sheinwoodian. The end is defined as the base (or GSSP A Global Boundary Stratotype Section and Point (GSSP), sometimes referred to as a golden spike, is an internationally agreed upo ...
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Ostracoderm
Ostracodermi () or ostracoderms is an informal group of vertebrate animals that include all armored jawless fish of the Paleozoic Era. The term does not often appear in classifications today because it is paraphyletic (excluding jawed fishes and possibly the cyclostomes if anaspids are closer to them) and thus does not correspond to one evolutionary lineage. However, the term is still used as an informal way of loosely grouping together the armored jawless fishes. An innovation of ostracoderms was the use of gills not for feeding, but exclusively for respiration. Earlier chordates with gill precursors used them for both respiration and feeding. Ostracoderms had separate pharyngeal gill pouches along the side of the head, which were permanently open with no protective operculum. Unlike invertebrates that use ciliated motion to move food, ostracoderms used their muscular pharynx to create a suction that pulled small and slow-moving prey into their mouths. Swiss anatomist L ...
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Silurian
The Silurian ( ) is a geologic period and system spanning 23.5 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the third and shortest period of the Paleozoic Era, and the third of twelve periods of the Phanerozoic Eon. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods ( myriapods, arachnids and hexapods) ...
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Order (biology)
Order () is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. For some groups of organisms, their orders may follow consist ...
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Psammosteida
Psammosteida also called as Psammosteoidei is a suborder of pteraspidid heterostracan agnathans. The psammosteids had broad, flattened bodies, suggesting a predominantly benthic habit. The earliest unequivocal psammosteid is '' Drepanaspis'' of Early Devonian Germany, which is either included in the family Psammosteidae, or placed within its own family, Drepanaspididae. If the late Silurian/Early Devonian '' Weigeltaspis'' is a psammosteid, as opposed to being a traquairaspid, then that genus, instead, would be the oldest psammosteid. However, its placement within Heterostraci remains a matter of debate. Other notable psammosteids include '' Psammosteus'', and '' Obruchevia'', two genera of enormous species with dorsal shields around one meter in diameter. The Psammosteids were the only heterostracans to survive to the end of the Devonian, where they finally perish during the Hangenberg event The Hangenberg event, also known as the Hangenberg crisis or end-Devonian ext ...
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Caudal Fin
Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported only by muscles. Fish fins are distinctive anatomical features with varying structures among different clades: in ray-finned fish (Actinopterygii), fins are mainly composed of bony spines or rays covered by a thin stretch of scaleless skin; in lobe-finned fish (Sarcopterygii) such as coelacanths and lungfish, fins are short rays based around a muscular central bud supported by jointed bones; in cartilaginous fish (Chondrichthyes) and jawless fish (Agnatha), fins are fleshy " flippers" supported by a cartilaginous skeleton. Fins at different locations of the fish body serve different purposes, and are divided into two groups: the midsagittal ''unpaired fins'' and the more laterally located ''paired fins''. Unpaired fins are predominan ...
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Thelodonti
Thelodonti (from Greek: "nipple teeth")Maisey, John G., Craig Chesek, and David Miller. Discovering fossil fishes. New York: Holt, 1996. is a class of extinct Palaeozoic jawless fishes with distinctive scales instead of large plates of armor. There is much debate over whether the group represents a monophyletic grouping, or disparate stem groups to the major lines of jawless and jawed fish. Thelodonts are united in possession of " thelodont scales". This defining character is not necessarily a result of shared ancestry, as it may have been evolved independently by different groups. Thus the thelodonts are generally thought to represent a polyphyletic group, although there is no firm agreement on this point. On the basis that they are monophyletic, they are reconstructed as being ancestrally marine and invading freshwater on multiple occasions. "Thelodonts" were morphologically very similar, and probably closely related, to fish of the classes Heterostraci and Anaspida, di ...
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Dentine
Dentin ( ) (American English) or dentine ( or ) (British English) () is a calcified tissue of the body and, along with enamel, cementum, and pulp, is one of the four major components of teeth. It is usually covered by enamel on the crown and cementum on the root and surrounds the entire pulp. By volume, 45% of dentin consists of the mineral hydroxyapatite, 33% is organic material, and 22% is water. Yellow in appearance, it greatly affects the color of a tooth due to the translucency of enamel. Dentin, which is less mineralized and less brittle than enamel, is necessary for the support of enamel. Dentin rates approximately 3 on the Mohs scale of mineral hardness. There are two main characteristics which distinguish dentin from enamel: firstly, dentin forms throughout life; secondly, dentin is sensitive and can become hypersensitive to changes in temperature due to the sensory function of odontoblasts, especially when enamel recedes and dentin channels become exposed. Devel ...
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Histology
Histology, also known as microscopic anatomy or microanatomy, is the branch of biology that studies the microscopic anatomy of biological tissue (biology), tissues. Histology is the microscopic counterpart to gross anatomy, which looks at larger structures visible without a microscope. Although one may divide microscopic anatomy into ''organology'', the study of organs, ''histology'', the study of tissues, and ''cytology'', the study of cell (biology), cells, modern usage places all of these topics under the field of histology. In medicine, histopathology is the branch of histology that includes the microscopic identification and study of diseased tissue. In the field of paleontology, the term paleohistology refers to the histology of fossil organisms. Biological tissues Animal tissue classification There are four basic types of animal tissues: muscle tissue, nervous tissue, connective tissue, and epithelial tissue. All animal tissues are considered to be subtypes of these ...
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Agnathan
Agnatha (; ) or jawless fish is a paraphyletic infraphylum of animals in the subphylum Vertebrata of the phylum Chordata, characterized by the lack of jaws. The group consists of both living ( cyclostomes such as hagfishes and lampreys) and extinct clades (e.g. conodonts and cephalaspidomorphs, among others). They are sister to vertebrates with jaws known as gnathostomes, who evolved from jawless ancestors during the early Silurian by developing folding articulations in the first pairs of gill arches. Molecular data, both from rRNA and from mtDNA as well as embryological data, strongly supports the hypothesis that both groups of living agnathans, hagfishes and lampreys, are more closely related to each other than to jawed fish, forming the superclass Cyclostomi. The oldest fossil agnathans appeared in the Cambrian. Living jawless fish comprise about 120 species in total. Hagfish are considered members of the subphylum Vertebrata, because they secondarily lost vertebr ...
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Devonian
The Devonian ( ) is a period (geology), geologic period and system (stratigraphy), system of the Paleozoic era (geology), era during the Phanerozoic eon (geology), eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago (Megaannum, Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of history of life#Colonization of land, life on land occurred during the Devonian, as free-spore, sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaf, leaves and true roots, and by the end of the period the first seed-bearing plants (Pteridospermatophyta, pteridospermatophyt ...
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Ordovician
The Ordovician ( ) is a geologic period and System (geology), system, the second of six periods of the Paleozoic Era (geology), Era, and the second of twelve periods of the Phanerozoic Eon (geology), Eon. The Ordovician spans 41.6 million years from the end of the Cambrian Period Megaannum, Ma (million years ago) to the start of the Silurian Period Ma. The Ordovician, named after the Celtic Britons, Welsh tribe of the Ordovices, was defined by Charles Lapworth in 1879 to resolve a dispute between followers of Adam Sedgwick and Roderick Murchison, who were placing the same Rock (geology), rock beds in North Wales in the Cambrian and Silurian systems, respectively. Lapworth recognized that the fossil fauna in the disputed Stratum, strata were different from those of either the Cambrian or the Silurian systems, and placed them in a system of their own. The Ordovician received international approval in 1960 (forty years after Lapworth's death), when it was adopted as an official per ...
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