Eureptiles
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Eureptiles
Sauropsida (Greek language, Greek for "lizard faces") is a clade of amniotes, broadly equivalent to the Class (biology), class Reptile, Reptilia, though typically used in a broader sense to also include extinct stem-group relatives of modern reptiles and birds (which, as theropod dinosaurs, are nested within reptiles as more closely related to crocodilians than to lizards or turtles).Gauthier J.A. (1994): ''The diversification of the amniotes''. In: D.R. Prothero and R.M. Schoch (ed.) Major Features of Vertebrate Evolution: 129–159. Knoxville, Tennessee: The Paleontological Society. The most popular definition states that Sauropsida is the Sister group, sibling taxon to Synapsida, the other clade of amniotes which includes mammals as its only modern representatives. Although early synapsids have historically been referred to as "mammal-like reptiles", all synapsids are more closely related to mammals than to any modern reptile. Sauropsids, on the other hand, include all amniotes m ...
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Pennsylvanian (geology)
The Pennsylvanian ( , also known as Upper Carboniferous or Late Carboniferous) is, on the International Commission on Stratigraphy, ICS geologic timescale, the younger of two period (geology), subperiods of the Carboniferous Period (or the upper of two system (stratigraphy), subsystems of the Carboniferous System). It lasted from roughly . As with most other geochronology, geochronologic units, the stratum, rock beds that define the Pennsylvanian are well identified, but the exact date of the start and end are uncertain by a few hundred thousand years. The Pennsylvanian is named after the U.S. state of Pennsylvania, where the coal Bed (geology), beds of this age are widespread. The division between Pennsylvanian and Mississippian (geology), Mississippian comes from North American stratigraphy. In North America, where the early Carboniferous beds are primarily marine limestones, the Pennsylvanian was in the past treated as a full-fledged geologic period between the Mississippian ...
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Parareptilia
Parareptilia ("near-reptiles") is an extinct group of basal sauropsids (" reptiles"), traditionally considered the sister taxon to Eureptilia (the group that likely contains all living reptiles and birds). Parareptiles first arose near the end of the Carboniferous period and achieved their highest diversity during the Permian period. Several ecological innovations were first accomplished by parareptiles among reptiles. These include the first reptiles to return to marine ecosystems (mesosaurs), the first bipedal reptiles ( bolosaurids such as '' Eudibamus''), the first reptiles with advanced hearing systems ( nycteroleterids and others), and the first large herbivorous reptiles (the pareiasaurs). The only parareptiles to survive into the Triassic period were the procolophonoids, a group of small generalists, omnivores, and herbivores. The largest family of procolophonoids, the procolophonids, rediversified in the Triassic, but subsequently declined and became extinct by t ...
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Thuringothyris
''Thuringothyris'' is an extinct genus of Early Permian eureptiles known from the Thuringian Forest in central Germany. Description ''Thuringothyris'' is known from the holotype MNG 7729, articulated well-preserved skull and partial postcranial skeleton, and from the referred specimens MNG 10652, poorly preserved skull and partial vertebral column, MNG 10647, disarticulated cranial and postcranial remains of at least four individuals, MNG 10183, slightly crushed skull and partial postcranial skeleton and MNG 11191, poorly preserved skull and partial limbs. All specimens were collected from the Tambach-Sandstein Member, the uppermost part of the Tambach Formation, dating to the Artinskian stage of the Late Cisuralian Series (or alternatively upper Rotliegend), about 284–279.5 million years ago. They were found in the Bromacker Quarry, the middle part of the Thuringian Forest, near the small town of Tambach-Dietharz. ''Thuringothyris'' was originally ...
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Hylonomus
''Hylonomus'' (; ''hylo-'' "forest" + ''nomos'' "dweller") is an extinct genus of reptile that lived during the Bashkirian stage of the Late Carboniferous. It is the earliest known crown group amniote and the oldest known unquestionable reptile, with the only known species being ''Hylonomus lyelli''. Despite being amongst the oldest known reptiles, it is not the most primitive member of the group, being a eureptile more derived than either parareptiles or captorhinids. Discovery and naming ''Hylonomus lyelli'' was first described by John William Dawson in 1860. The species' name was given in honor of Dawson's teacher, the geologist Sir Charles Lyell. While it has traditionally been included in the group Protorothyrididae, it has since been recovered outside this group. Formerly assigned species Dawson also attributed two other species ''H. aciedentatus'' and ''H. wymani'' when he described ''H. lyelli'' in 1860, and later described two more species ''H. multidens'' and ''H. ...
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Coelostegus
''Coelostegus'' is an extinct genus of Late Carboniferous (late Westphalian stage) basal reptile known from Plzeň of Czech Republic. It is known from the holotype ČGH 3027, a partial skeleton of an immature individual. It was collected in the Nýřany site from the Nýřany Member of the Kladno Formation. It was first named by Robert L. Carroll and Donald Baird in 1972 and the type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ... is ''Coelostegus prothales''. The most recent phylogenic study of primitive reptile relationships found ''Coelostegus'' to be the basalmost known eureptile. References Protorothyrididae Carboniferous reptiles of Europe Fossil taxa described in 1972 {{paleo-reptile-stub ...
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Captorhinidae
Captorhinidae is an extinct family of tetrapods, traditionally considered primitive Reptile, reptiles, known from the late Carboniferous to the Late Permian. They had a cosmopolitan distribution across Pangea. Description Captorhinids are a clade of small to very large lizard-like animals that date from the Late Carboniferous through the Permian. Their skulls were much stronger than those of their relatives, the protorothyridids, and had teeth that were better able to deal with tough plant material. The postcranial skeleton is similar to those of seymouriamorphs and diadectomorphs; these animals were grouped together with the captorhinids in the order Cotylosauria as the first reptiles in the early 20th century, but are now usually regarded as Stem group, stem-amniotes no closer to reptiles than to mammals. Captorhinids have broad, robust skulls that are generally triangular in shape when seen in dorsal view. The premaxillae are characteristically downturned. The largest captorh ...
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Procolophonia
Procolophonia is an extinct suborder (clade) of herbivorous reptiles that lived from the Middle Permian till the end of the Triassic period. They were originally included as a suborder of the Cotylosauria (later renamed Captorhinida Carroll 1988) but are now considered a clade of Parareptilia. They are closely related to other generally lizard-like Permian reptiles such as the Millerettidae, Bolosauridae, Acleistorhinidae, and Lanthanosuchidae, all of which are included under the Anapsida or "Parareptiles" (as opposed to the Eureptilia). Classification There are two main groups of Procolophonia, the small, lizard-like Procolophonoidea, and the Pareiasauroidea, which include the large, armoured Pareiasauridae. According to the traditional classification of Carroll 1988 as well as phylogenetic analyses of 2012, smaller groups like Rhipaeosauridae (now a synonym of Nycteroleteridae) and Sclerosauridae are classified with the pareiasaurs and with the procolophonids, ...
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Acleistorhinidae
Acleistorhinidae is an extinct family of Late Carboniferous and Early Permian-aged ( Moscovian to Kungurian stage) parareptiles. It is defined as a node based clade including the last common ancestor of '' Acleistorhinus pteroticus'' and '' Colobomycter pholeter'' and all its descendants. Acleistorhinids are most diverse from the Richards Spur locality of the Early Permian of Oklahoma. Richards Spur acleistorhinids include '' Acleistorhinus'', '' Colobomycter'', '' Delorhynchus'', '' Feeserpeton'' and '' Klastomycter''. Other taxa include '' Carbonodraco'' from the Late Carboniferous of Ohio and '' Karutia'' from the Early Permian of Brazil. Acleistorhinidae is commonly considered a subgroup of lanthanosuchoids, related to taxa such as '' Chalcosaurus'', '' Lanthaniscus'' and '' Lanthanosuchus''. However, a re-examination of parareptile phylogeny conducted by Cisneros ''et al.'' (2021) argued that lanthanosuchids were not closely related to acleistorhinids. The phylogenetic ...
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Bolosauria
Procolophonomorpha is an Order (biology), order containing most parareptiles. Many papers have applied various definitions to the name, though most of these definitions have since been considered synonymous with modern parareptile Clade, clades such as Ankyramorpha and Procolophonia. The current definition of Procolophonomorpha, as defined by Modesto, Scott, & Reisz (2009), is that of as a stem-based group containing ''Procolophon'' and all taxa more closely related to it than to ''Milleretta''. It constitutes a diverse assemblage that includes a number of lizard-like forms, as well as more diverse types such as the pareiasaurs. Lee 1995, 1996, 1997 argues that turtles evolved from pareiasaurs, but this view is no longer considered likely. Rieppel and deBraga 1996 and deBraga and Rieppel, 1997 argue that turtles evolved from sauropterygians, and there is both molecular and fossil (''Pappochelys'') evidence for the origin of turtles among diapsid reptiles. Classification The foll ...
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Millerettidae
Millerettidae is an extinct family of parareptiles from the Middle Permian to the Late Permian period ( Capitanian - Changhsingian stages) of South Africa. The millerettids were small insectivores and probably resembled modern lizards in appearance and lifestyle. The following cladogram shows the phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ... position of the Millerettidae, from Ruta ''et al.'', 2011. References External linksBerkeley University information. Permian reptiles Prehistoric reptiles of Africa Guadalupian first appearances Lopingian extinctions Prehistoric reptile families {{Permian-reptile-stub ...
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Mesosauria
Mesosaurs ("middle lizards") were a group of small aquatic reptiles that lived during the early Permian period (Cisuralian), roughly Geologic time scale, 299 to 270 million years ago. Mesosaurs were the first known aquatic reptiles, having apparently returned to an aquatic lifestyle from more terrestrial ancestors. It is uncertain which and how many terrestrial traits these ancestors displayed; recent research cannot establish with confidence if the first amniotes were fully terrestrial, or only amphibious. Most authors consider mesosaurs to have been aquatic, although adult animals may have been amphibious, rather than completely aquatic, as indicated by their moderate skeletal adaptations to a semiaquatic lifestyle.Pablo Nuñez Demarco et al. Was Mesosaurus a Fully Aquatic Reptile? Front. Ecol. Evol, published online July 27, 2018; doi: 10.3389/fevo.2018.00109 Similarly, their affinities are uncertain; they may have been among the most basal sauropsids or among the most basal p ...
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Microleter
''Microleter'' is an extinct genus of basal (phylogenetics), basal Procolophonomorpha, procolophonomorph parareptiles which lived in Oklahoma during the Early Permian period. The type species, type and only known species is ''Microleter mckinzieorum''. ''Microleter'' is one of several parareptile taxa described from the Richards Spur fissure fills, and can be characterized from its high tooth count, Lacrimal bone, lacrimal/Nostril, narial contact, short Postfrontal bone, postfrontal, and slit-like Infratemporal fenestra, temporal emargination edged by the Postorbital bone, postorbital, Jugal bone, jugal, Squamosal bone, squamosal, and Quadratojugal bone, quadratojugal. Contrary to ''Australothyris'', which had a similar Phylogenetics, phylogenetic position as a basal procolophonomorph, ''Microleter'' suggests that early parareptile evolution occurred in Laurasia and that multiple lineages developed openings or emarginations in the temporal region. Discovery The only known specim ...
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